
Scheiner, S. M. & Willig, M. R. A general theory of ecology. Theoretical Ecology 1, 21–28 (2008).
Collins, C. & Kays, R. Causes of mortality in North American populations of large and medium‐sized mammals. Animal Conservation 14, 474–483 (2011).
Hill, J., DeVault, T. L. & Belant, J. L. Cause–specific mortality of the world’s terrestrial vertebrates. Global Ecology and Biogeography 28, 680–689 (2019).
Gibbs, J. P. & Steen, D. A. Trends in sex ratios of turtles in the United States: implications of road mortality. Conservation Biology 19, 552–556 (2005).
Wright, G. J., Peterson, R. O., Smith, D. W. & Lemke, T. O. Selection of northern Yellowstone elk by gray wolves and hunters. Journal of Wildlife Management 70, 1070–1078 (2006).
Kays, R. et al. Does hunting or hiking affect wildlife communities in protected areas? Journal of Applied Ecology 54, 242–252 (2017).
Leclerc, M., Frank, S. C., Zedrosser, A., Swenson, J. E. & Pelletier, F. Hunting promotes spatial reorganization and sexually selected infanticide. Scientific Reports 7, 1–6 (2017).
Hill, J., DeVault, T. L. & Belant, J. L. Anthropogenic mammal mortality increases with the human footprint. Frontiers in Ecology and the Environment 18, 13–18 (2019).
Milner, J. M., Nilsen, E. B. & Reassen, H. P. Demographic side effects of selective hunting in ungulates and carnivores. Conservation Biology 21, 36–47 (2006).
Kristensen, T., Means, M., Eggert, L. S., Smith, K. G. & White, D. Demographics of American black bear populations following changes in harvest policy. Ursus 29, 147–162 (2019).
Ripple, W. J. et al. Status and ecological effects of the world’s largest carnivores. Science 343, 1241484 (2014).
Darimont, C. T., Fox, C. H., Bryan, H. M. & Reimchen, T. E. The unique ecology of human predators. Science 349, 858–860 (2015).
Krofel, M., Treves, A., Ripple, W. J., Chapron, G. & López-Bao, J. V. Hunted carnivores at outsized risk. Science 350, 518–519 (2015).
Festa-Bianchet, M. & Mysterud, A. Hunting and evolution: theory, evidence, and unknowns. Journal of Mammalogy 99, 1281–1292 (2018).
Brown, J. H. & Nicoletto, P. F. Spatial scaling of species composition: body masses of North American land mammals. The American Naturalist 138, 1478–1512 (1991).
Mcloughlin, P. D., Ferguson, S. H. & Messier, F. Intraspecific variation in home range overlap with habitat quality: a comparison among brown bear populations. Evolutionary Ecology 14, 39–60 (2000).
Gantchoff, M., Wang, G., Beyer, D. & Belant, J. L. Scale‐dependent home range optimality for a solitary omnivore. Ecology and Evolution 8, 12271–12282 (2018).
Schwab, A. C. & Zandbergen, P. A. Vehicle–related mortality and road crossing behavior of the Florida panther. Applied Geography 31, 859–870 (2011).
Treves, A. & Karanth, K. U. Human‐carnivore conflict and perspectives on carnivore management worldwide. Conservation Biology 17, 1491–1499 (2003).
Beckmann, J. P. & Berger, J. Rapid ecological and behavioural changes in carnivores: the responses of black bears (Ursus americanus) to altered food. Journal of Zoology 261, 207–212 (2003).
Feng, Y. et al. Bear bile: dilemma of traditional medicinal use and animal protection. Journal of Ethnobiology and Ethnomedicine 5, 2 (2009).
Albert, D. M., Bowyer, R. T. & Miller, S. D. Effort and success of brown bear hunters in Alaska. Wildlife Society Bulletin 29, 501–508 (2001).
Freeman, M. M. & Wenzel, G. W. The nature and significance of polar bear conservation hunting in the Canadian. Arctic. Arctic 59, 21–30 (2006).
Beston, J. A. Variation in life history and demography of the American black bear. The Journal of Wildlife Management 75, 1588–1596 (2011).
Freedman, A. H., Portier, K. M. & Sunquist, M. E. Life history analysis for black bears (Ursus americanus) in a changing demographic landscape. Ecological Modelling 167, 47–64 (2003).
Hebblewhite, M., Percy, M. & Serrouya, R. Black bear (Ursus americanus) survival and demography in the Bow Valley of Banff National Park, Alberta. Biological Conservation 112, 415–425 (2003).
Scheick, B. K. & McCown, W. Geographic distribution of American black bears in North America. Ursus 25, 24–34 (2014).
Garshelis, D. L., Scheick, B. K., Doan–Crider, D. L., Beecham, J. J. & Obbard, M. E. Ursus americanus. The IUCN Red List of Threatened Species 2016, e.T41687A45034604 (2016).
Bales, S. L., Hellgren, E. C., Leslie, D. M. Jr. & Hemphill, J. Jr. Dynamics of a recolonizing population of black bears in the Ouachita Mountains of Oklahoma. Wildlife Society Bulletin 33, 1342–1351 (2005).
Simek, S. L., Belant, J. L., Young, B. W., Shropshire, C. & Leopold, B. D. History and status of the American black bear in Mississippi. Ursus 23, 159–167 (2012).
Clark, J. D., Huber, D. & Servheen, C. Bear reintroductions: lessons and challenges. Ursus 13, 335–345 (2002).
Larkin, J. L., Cox, J. J., Wichrowski, M. W., Dzialak, M. R. & Maehr, D. S. Influences on release‐site fidelity of translocated elk. Restoration Ecology 12, 97–105 (2004).
Fritts, S. H. et al. Planning and implementing a reintroduction of wolves to Yellowstone National Park and central Idaho. Restoration Ecology 5, 7–27 (1997).
Belant, J. L., Etter, D. R., Friedrich, P., Williams, B. & Scribner, K. Comparison of techniques for sex determination of American martens. Journal of Wildlife Management 75, 256–260 (2011).
Norton, D. C., Belant, J. L., Beyer, D. E. Jr. & Bruggink, J. G. Female American black bears do not alter space use or movements to reduce infanticide risk. Plos One. 13, e0203651 (2018).
Wilton, C. M. et al. Trap array configuration influences estimates and precision of black bear density and abundance. PLoS One 9, e111257 (2014).
Duquette, J. F. et al. Black bear (Ursus americanus) functional resource selection relative to intraspecific competition and human risk. Canadian Journal of Zoology 95, 203–212 (2017).
Kays, R., Crofoot, M. C., Jetz, W. & Wikelski, M. Terrestrial animal tracking as an eye on life and planet. Science 348, aaa2478 (2015).
Naef‐Daenzer, B., Korner‐Nievergelt, F., Fiedler, W. & Grüebler, M. U. Bias in ring‐recovery studies: causes of mortality of little owls Athene noctua and implications for population assessment. Journal of Avian Biology 48, 266–274 (2017).
Gantchoff, M., Conlee, L. & Belant, J. L. Conservation implications of sex‐specific landscape suitability for a large generalist carnivore. Diversity and Distributions 25, 1488–1496 (2019).
Lunn, M. & McNeil, D. Applying Cox regression to competing risks. Biometrics 51, 524–532 (1995).
Murray, D. L. et al. Death from anthropogenic causes is partially compensatory in recovering wolf populations. Biological Conservation 143, 2514–2524 (2010).
Grambsch, P. & Therneau, T. Proportional hazards tests and diagnostics based on weighted residuals. Biometrika 81, 515–26 (1994).
R Core Team R: A language and environment for statistical computing. R Foundation for Statistical Computing, Vienna, Austria. Available from, https://www.R–project.org/ (2019).
Therneau, T. M. & Lumley, T. Survival: survival analyses. R package version 2.44–1.1. Available from, https://cran.r–project.org/web/packages/survival/index.html (2019).
Bled, F. & Belant, J. L. demetR: a Bayesian population simulation web–application for harvest management. Ursus 29, 82–92 (2019).
Hill, J., DeVault, T. L. & Belant, J. L. CauseSpec: A database of global terrestrial vertebrate mortality. Ecology, e02865, https://doi.org/10.1002/ecy.2865 (2019).
Warton, D. I. & Hui, F. K. The arcsine is asinine: the analysis of proportions in ecology. Ecology 92, 3–10 (2011).
Burnham, K. P. & Anderson, D. R. Model selection and multimodel inference: a practical information–theoretical approach. (Springer, 2002).
Bischof, R., Fujita, R., Zedrosser, A., Söderberg, A. & Swenson, J. E. Hunting patterns, ban on baiting, and harvest demographics of brown bears in Sweden. The Journal of Wildlife Management 72, 79–88 (2008).
Cooley, H. S., Wielgus, R. B., Koehler, G. M., Robinson, H. S. & Maletzke, B. T. Does hunting regulate cougar populations? A test of the compensatory mortality hypothesis. Ecology 90, 2913–2921 (2009).
Keller, B. J. et al. A review of vital rates and cause‐specific mortality of elk Cervus elaphus populations in eastern North America. Mammal Review 45, 146–159 (2015).
Bischof, R., Swenson, J. E., Yoccoz, N. G., Mysterud, A. & Gimenez, O. The magnitude and selectivity of natural and multiple anthropogenic mortality causes in hunted brown bears. Journal of Animal Ecology 78, 656–665 (2009).
Obbard, M. E. et al. Big enough for bears? American black bears at heightened risk of mortality during seasonal forays outside Algonquin Provincial Park, Ontario. Ursus 28, 182–195 (2017).
Pigeon, G., Festa‐Bianchet, M., Coltman, D. W. & Pelletier, F. Intense selective hunting leads to artificial evolution in horn size. Evolutionary Applications 9, 521–530 (2016).
McLellan, B. N. & Shackleton, D. M. A comparison of grizzly bear harvest data from Montana and southeastern British Columbia. Wildlife Society Bulletin 16, 371–375 (1988).
Czetwertynski, S. M., Boyce, M. S. & Schmiegelow, F. K. Effects of hunting on demographic parameters of American black bears. Ursus 18, 1–18 (2007).
Regehr, E. V., Wilson, R. R., Rode, K. D., Runge, M. C. & Stern, H. L. Harvesting wildlife affected by climate change: a modelling and management approach for polar bears. Journal of Applied Ecology 54, 1534–1543 (2017).
Rughetti, M. & Festa‐Bianchet, M. Effects of selective harvest of non‐lactating females on chamois population dynamics. Journal of Applied Ecology 51, 1075–1084 (2014).
Krofel, M., Jonozovič, M. & Jerina, K. Demography and mortality patterns of removed brown bears in a heavily exploited population. Ursus 23, 91–103 (2012).
Van de Walle, J., Pigeon, G., Zedrosser, A., Swenson, J. E. & Pelletier, F. Hunting regulation favors slow life histories in a large carnivore. Nature Communications 9, 1100 (2018).
Gantchoff, M. G., Beyer, D. & Belant, J. L. Reproductive class influences risk tolerance during denning and spring for American black bears (Ursus americanus). Ecosphere 10, e02705 (2019).
Coltman, D. W. et al. Undesirable evolutionary consequences of trophy hunting. Nature 426, 655–658 (2003).
Di Minin, E., Leader-Williams, N. & Bradshaw, C. J. Banning trophy hunting will exacerbate biodiversity loss. Trends in Ecology and Evolution 31, 99–102 (2016).
Wanger, T. C., Traill, L. W., Cooney, R., Rhodes, J. R. & Tscharntke, T. Trophy hunting certification. Nature Ecology and Evolution 1, 1791–1793 (2017).
Bischof, R. et al. Implementation uncertainty when using recreational hunting to manage carnivores. Journal of Applied Ecology 49, 824–832 (2012).
Olson, E. R. et al. Pendulum swings in wolf management led to conflict, illegal kills, and a legislated wolf hunt. Conservation Letters 8, 351–360 (2015).
Mahoney, S. P. Recreational hunting and sustainable wildlife use in North America. In Dickson, B., Hutton, J. & Adams, W.M. (eds.), Recreational hunting, conservation and rural livelihoods: Science and practice. (Wiley-Blackwell, 2009).
Jacobson, C. A., Organ, J. F., Decker, D. J., Batcheller, G. R. & Carpenter, L. A conservation institution for the 21st century: Implications for state wildlife agencies. The Journal of Wildlife Management 74, 203–209 (2010).
IUCN IUCN SSC Guiding principles on trophy hunting as a tool for creating conservation incentives. International Union for the Conservation of Nature. Gland, Switzerland, Ver. 1.0 (2012).
Carter, N. H. et al. A conceptual framework for understanding non-harvest anthropogenicing of large carnivores. Ambio 46, 251–264 (2017).
Miller, S. D. & Tutterrow, V. L. Characteristics of nonsport mortalities to brown and black bears and human injuries from bears in Alaska. Ursus 11, 239–252 (1999).
Liberg, O. et al. Shoot, shovel and shut up: Cryptic poaching slows restoration of a large carnivore in Europe. Proceedings of the Royal Society of London, Series B: Biological Sciences 270, 91–98 (2012).
Heurich, M. et al. Illegal hunting as a major driver of the source-sink dynamics of a reintroduced lynx population in Central Europe. Biological Conservation 224, 355–365 (2018).
Červený, J., Krojerová-Prokešová, J., Kušta, T. & Koubek, P. The change in the attitudes of Czech hunters towards Eurasian lynx: Is poaching restricting lynx population growth? Journal for Nature Conservation 47, 28–37 (2019).
Kaczensky, P. et al. Non-harvest anthropogenicings may hamper brown bear recovery in the Eastern Alps. Ursus 22, 37–46 (2011).
McCarthy, T. M. & Seavoy, R. J. Reducing nonsport losses attributable to food conditioning: human and bear behavior modification in an urban environment. Bears: Their Biology and Management 9, 75–84 (1994).
Siemer, W. F., Hart, P. S., Decker, D. J. & Shanahan, J. E. Factors that influence concern about human–black bear interactions in residential settings. Human Dimensions of Wildlife 14, 185–197 (2009).
Creel, S. & Rotella, J. J. Meta–analysis of relationships between human offtake, total mortality and population dynamics of gray wolves (Canis lupus). PLoS ONE 5, 1–7 (2010).
Wolfe, M. L. et al. Is anthropogenic cougar mortality compensated by changes in natural mortality in Utah? Insight from long–term studies. Biological Conservation 182, 187–196 (2015).
Rice, M. B., Ballard, W. B., Fish, E. B., McIntyre, N. E. & Holdermann, D. Habitat–distribution modeling of a recolonizing black bear, Ursus americanus, population in the Trans–Pecos region of Texas. The Canadian Field–Naturalist 123, 246–254 (2009).
Smith, J. A., Duane, T. P. & Wilmers, C. C. Moving through the matrix: Promoting permeability for large carnivores in a human–dominated landscape. Landscape and Urban Planning 183, 50–58 (2019).
Woodroffe, R. & Redpath, S. M. When the hunter becomes the hunted. Science 348, 1312–1314 (2015).
Mitchell, B. R., Jaeger, M. M. & Barrett, R. H. Coyote depredation management: current methods and research needs. Wildlife Society Bulletin 32, 1209–1218 (2004).
Hody, J. W. & Kays, R. Mapping the expansion of coyotes (Canis latrans) across North and Central America. ZooKeys 759, 81–97 (2018).
Stillfried, M., Belant, J. L., Svoboda, N. J., Beyer, D. E. & Kramer-Schadt, S. When top predators become prey: black bears alter movement behaviour in response to hunting pressure. Behavioural Processes 120, 30–39 (2015).
Ditmer, M. A., Noyce, K. V., Fieberg, J. R. & Garshelis, D. L. Delineating the ecological and geographic edge of an opportunist: The American black bear exploiting an agricultural landscape. Ecological Modelling 387, 205–219 (2018).
Baruch-Mordo, S. et al. Stochasticity in natural forage production affects use of urban areas by black bears: implications to management of human-bear conflicts. PloS ONE 9, e85122 (2014).
Treves, A., Artelle, K. A., Darimont, C. T. & Parsons, D. R. Mismeasured mortality: correcting estimates of wolf poaching in the United States. Journal of Mammalogy 98, 1256–1264 (2017).
Koons, D. N. et al. Methods for studying cause‐specific senescence in the wild. Methods in Ecology and Evolution 5, 924–933 (2014).
Bischof, R. et al. Regulated hunting re-shapes the life history of brown bears. Nature Ecology and Evolution 2, 116–123 (2018).
Hill, J., DeVault, T. L. & Belant, J. L. Protected areas reduce poaching but not overall anthropogenic mortality of North American mammals. Global Ecology and Conservation 21, e00810 (2019).
Baeza, A. & Estades, C. F. Effect of the landscape context on the density and persistence of a predator population in a protected area subject to environmental variability. Biological Conservation 143, 94–101 (2010).
Johnson, A., Vongkhamheng, C., Hedemark, M. & Saithongdam, T. Effects of human–carnivore conflict on tiger (Panthera tigris) and prey populations in Lao PDR. Animal Conservation 9, 421–430 (2006).
Carter, N. H. & Linnell, J. D. Co–adaptation is key to coexisting with large carnivores. Trends in Ecology and Evolution 31, 575–578 (2016).
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