
Mahler, A. E. Activity budgets and use of exhibit space by South American tapir (Tapirus terrestris) in a Zoological Park setting. Zoo. Biol. 3, 35–46 (1984).
Read, B. Breeding and management of the Malayan tapir: at St Louis Zoo. Int. Zoo. Yearbook. 24, 294–297 (1986).
Barongi, R. A. Husbandry and conservation of tapirs Tapirus spp. Int. Zoo. Yearbook. 32, 7–15 (1993).
Williams, K. D. & Petrides, G. A. Browse use, feeding behavior and management of Malayan Tapir. J. Wildl. Manag. 44, 489–494 (1980).
Traeholt, C. et al. Tapirus indicus. The IUCN Red List of Threatened Species 2016: e.T21472A45173636. https://doi.org/10.2305/IUCN.UK.2016-1.RLTS.T21472A45173636.en (2016).
Pukazhenthi, B. et al. A review of the reproductive biology and breeding management of tapirs. Integr. zool. 8, 18–34 (2013).
Lilia, K., Rosnina, Y., Abd Wahid, H., Zahari, Z. Z. & Abraham, M. Gross anatomy and ultrasonographic images of the reproductive system of the Malayan Tapir (Tapirus indicus). Anat. Histol. Embryol. 39, 569–575 (2010).
Arumugam, K. A. et al. Influence of enclosure conditions and visitors on the behavior of captive Malayan tapir (Tapirus indicus): implications for ex-situ management and conservation. IJSRP. 8, 22–33 (2018).
Tortato, M. A., Oliveira-Santos, L. G. R., Machado Filho, L. C. P., Brusius, L. & Hotzel, M. J. Reproductive behaviour repertoire of semi-captive lowland tapir Tapirus terrestris (Linnaeus, 1758). Biotemas. 20, 135–139 (2007).
Amoah, E. A., Gelaye, S., Guthrie, P. & Rexroad, C. E. Breeding season and aspects of reproduction of female goats. J. Anim. Sci. 74, 723–728 (1996).
Bronson, F. H. Climate change and seasonal reproduction in mammals. Philos. Trans. R. Soc. Lond. B Biol. Sci. 364, 3331–3340 (2009).
Mysterud, A., Langvatn, R. & Stenseth, N. C. Patterns of reproductive effort in male ungulates. J. Zool. 264, 209–215 (2004).
Aurich, C. Reproductive cycles of horses. Anim. Reprod. Sci. 124, 220–228 (2011).
Bercovitch, F. B., Bashaw, M. J. & del Castillo, S. M. Sociosexual behavior, male mating tactics, and the reproductive cycle of giraffe Giraffa camelopardalis. Horm. Behav. 50, 314–321 (2006).
Brown, J. L. et al. Reproductive activity in captive female cheetahs (Acinonyx jubatus) assessed by faecal steroids. Reproduction. 106, 337–346 (1996).
Zhang, G., Swaisgood, R. R. & Zhang, H. Evaluation of behavioral factors influencing reproductive success and failure in captive giant pandas. Zoo. Biol. 23, 15–31 (2004).
Carlstead, K., Fraser, J., Bennett, C. & Kleiman, D. G. Black rhinoceros (Diceros bicornis) in U.S. zoos: II. behavior, breeding success, and mortality in relation to housing facilities. Zoo. Biol. 18, 35–52 (1999).
Quadros, S., Goulart, V. D., Passos, L., Vecci, M. A. & Young, R. J. Zoo visitor effect on mammal behaviour: Does noise matter? Appl. Anim. Behav. Sci. 156, 78–84 (2014).
Hosey, G. R. Zoo animals and their human audiences: what is the visitor effect? Anim. Welf. 9, 343–357 (2000).
Morgan, K. N. & Tromborg, C. T. Sources of stress in captivity. Appl. Anim. Behav. Sci. 102, 262–302 (2007).
Larsen, M. J., Sherwen, S. L. & Rault, J. L. Number of nearby visitors and noise level affect vigilance in captive koalas. Appl. Anim. Behav. Sci. 154, 76–82 (2014).
Rajagopal, T., Archunan, G. & Sekar, M. Impact of zoo visitors on the fecal cortisol levels and behavior of an endangered species: Indian blackbuck (Antelope cervicapra L.). J. Appl. Anim. Welf. Sci. 14, 18–32 (2011).
Owen, M. A., Swaisgood, R. R., Czekala, N. M., Steinman, K. & Lindburg, D. G. Monitoring stress in captive giant pandas (Ailuropoda melanoleuca): behavioral and hormonal responses to ambient noise. Zoo. Biol. 23, 147–164 (2004).
Liu, J. et al. Stereotypic behavior and fecal cortisol level in captive giant pandas in relation to environmental enrichment. Zoo. Biol. 25, 445–459 (2006).
Carlstead, K. & Brown, J. L. Relationships between patterns of fecal corticoid excretion and behavior, reproduction, and environmental factors in captive black (Diceros bicornis) and white (Ceratotherium simum) rhinoceros. Zoo. Bio. 24, 215–232 (2005).
Grau, G. A. Olfaction and reproduction in ungulates. Mammalian olfaction, Reprod. processes, behavior. 219, 242 (1976).
AZA. Tapir (Tapiridae) Care Manual. Association of Zoos and Aquariums, Silver Spring, Maryland: Association of Zoos and Aquariums. 65 (2013).
Gilmore, M. Tapir behaviour: An examination of activity patterns, mother young interactions, spatial use, and environmental effects in captivity on two species (Tapirus indicus and Tapirus bairdii). Masters Thesis. Oklahoma State University, USA (2007).
McDonnell, S. M. & Haviland, J. C. S. Agonistic ethogram of the equid bachelor band. Appl. Anim. Behav. Sci. 43, 147–188 (1995).
Metrione, L.C. Relationships of social behaviour and the captive environment to reproduction in female southern white rhinoceros (Ceratotherium simum simum). PhD Thesis. The Ohio State University, USA (2010).
Naundrup, P. J. Vocal behaviour and communication of the Malayan tapir (Tapirus indicus). Tapir Conservation. 21, 8–13 (2012).
Zahari, Z. Z., Rosnina, Y., Wahid, H., Yap, K. C. & Jainudeen, M. R. Reproductive behaviour of captive Sumatran rhinoceros. Anim. Reprod. Sci. 85, 327–335 (2005).
Martin P. & Bateson P. Measuring Behaviour: An Introductory Guide, second ed. Cambridge University Press. 176 (2007).
Gelman, A. Scaling regression inputs by dividing by two standard deviations. Stat. med. 27, 2865–2873 (2008).
Burnham, K. P. & Anderson, D. R. Model selection and multimodel inference. Springer Verlag (2002).
Bartoń, K. Multi-model inference. R package version 1.15.6 (2016).
Burnham, K. P., Anderson, D. R. & Huyvaert, K. P. AIC model selection and multimodel inference in behavioral ecology: some background, observations, and comparisons. Behav. Ecol. Sociobiol. 65, 23–35 (2011).
Campbell, C. J. Patterns of behavior across reproductive states of free–ranging female black–handed spider monkeys (Ateles geoffroyi). Am. J. Phys. Anthropol. 124, 166–176 (2004).
Taylor, S. A., Salo, A. L. & Dewsbury, D. A. Estrus induction in four species of voles (Microtus). J. Comp. Psychol. 106, 366 (1992).
Roelofs, J. B., Van Eerdenburg, F. J. C. M., Soede, N. M. & Kemp, B. Various behavioral signs of estrous and their relationship with time of ovulation in dairy cattle. Theriogenology. 63, 1366–1377 (2005).
Asa, C. S., Goldfoot, D. A. & Ginther, O. J. Assessment of the sexual behavior of pregnant mares. Horm. Behav. 17, 405–413 (1983).
Crowell-Davis, S. L. Sexual behaviour of mares. Horm. Behav. 52, 12–17 (2007).
Luna-Orozco, J. R., Fernández, I. G., Gelez, H. & Delgadillo, J. A. Parity of female goats does not influence their estrous and ovulatory responses to the male effect. Anim. Reprod. Sci. 106, 352–360 (2008).
Kretzschmar, P., Gansloßer, U. & Dehnhard, M. Relationship between androgens, environmental factors and reproductive behavior in male white rhinoceros (Ceratotherium simum simum). Horm. Behav. 45, 1–9 (2004).
Holden, J., Yanuar, A. & Martyr, D. J. The Asian Tapir in Kerinci Seblat National Park, Sumatra: evidence collected through photo-trapping. Oryx. 37, 34–40 (2003).
Brown, R. E., & Macdonald, D. W. Social odours in mammals. Clarendon Press (1985).
Brockman, D. K. Reproductive behavior of female Propithecus verreauxi at Beza Mahafaly, Madagascar. Int. J. Primat. 20, 375–398 (1999).
Hermes, R. et al. The effect of long non-reproductive periods on the genital health in captive female white rhinoceroses (Ceratotherium simum simum, C.s. cottoni). Theriogenology. 65, 1492–1515 (2006).
Shrestha, T. K. Mammals of Nepal (1997).
Hazarika, B. C. & Saikia, P. K. A study on the behaviour of Great Indian One-horned Rhino (Rhinoceros unicornis Linn.) in the Rajiv Gandhi Orang National Park, Assam, India. NeBIO. 1 (2010).
Roman, J. International Studbook Central American Tapir. Virginia Zoological Park. 104 (2009).
Roman, J. International Baird’s Tapir Studbook. Virginia Zoological Park. (2010).
Kusuda, S. et al. Estrous cycle based on blood progesterone profiles and changes in vulvar appearance of Malayan Tapirs (Tapirus indicus). J. Reprod. Develop. 53, 1283–1289 (2007).
Watts, J. M. & Stookey, J. M. Vocal behaviour in cattle: the animal’s commentary on its biological processes and welfare. Appl. Anim. Behav. Sci. 67, 15–33 (2000).
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