Bourke, A. F. G. & Heinze, J. The ecology of communal breeding: The case of multiple-queen Leptothoracine ants. Philos. Trans. R. Soc. Lond. B Biol. Sci.345, 359–372 (1994).
Bourke, A. F. G. Principles of Social Evolution. Oxford Series in Ecology and Evolution (2011).
Cockburn, A. Evolution of helping behavior in cooperatively breeding birds. Annu. Rev. Ecol. Evol. Syst.29, 141–177 (1998).
Jennions, M. Cooperative breeding in mammals. Trends Ecol. Evol.9, 89–93 (1994).
Lukas, D. & Clutton-Brock, T. Life histories and the evolution of cooperative breeding in mammals. Proc. R. Soc. B279, 4065–4070 (2012).
Purcell, J. Geographic patterns in the distribution of social systems in terrestrial arthropods. Biol. Rev.86, 475–491 (2011).
Wong, M. & Balshine, S. The evolution of cooperative breeding in the African cichlid fish, Neolamprologus pulcher. Biol. Rev.86, 511–530 (2011).
Dugatkin, L. Animal cooperation among unrelated individuals. Naturwissenschaften89, 533–541 (2002).
Emlen, S. The evolution of helping. An ecological constraints model. Am. Nat.119, 29–39 (1982).
Nichols, H. J. et al. Food availability shapes patterns of helping effort in a cooperative mongoose. Anim. Behav.83, 1377–1385 (2012).
Riehl, C. & Strong, M. J. Stable social relationships between unrelated females increase individual fitness in a cooperative bird. Proc. R. Soc. B285, 20180130 (2018).
Sharp, S. P., English, S. & Clutton-Brock, T. H. Maternal investment during pregnancy in wild meerkats. Evol. Ecol.27, 1033–1044 (2012).
Taborsky, M. Broodcare helpers in the cichlid fish Lamprologus brichardi: Their costs and benefits. Anim. Behav.32, 1236–1252 (1984).
Hamilton, W. D. The genetical evolution of social behaviour. J. Theor. Biol.7, 1–52 (1964).
Bshary, R. Cooperation between unrelated individuals—a game theoretic approach. In Animal Behaviour: Evolution and Mechanisms (ed. Kappeler, P.) 213–240 (Springer, Berlin, 2010).
Dugatkin, L. A. & Mesterton-Gibbons, M. Cooperation among unrelated individuals: Reciprocal altruism, by-product mutualism and group selection in fishes. Biosystems37, 19–30 (1996).
Keller, L. Queen Number and Sociality in Insects (Oxford University Press, Oxford, 1993).
Matsuura, K., Fujimoto, M., Goka, K. & Nishida, T. Cooperative colony foundation by termite female pairs: Altruism for survivorship in incipient colonies. Anim. Behav.64, 167–173 (2002).
Mesterton-Gibbons, M. & Dugatkin, L. A. Cooperation among unrelated individuals: Evolutionary factors. Q. Rev. Biol.67, 267–281 (1992).
Bernasconi, G. & Strassmann, J. E. Cooperation among unrelated individuals: The ant foundress case. Trends Ecol. Evol.14, 477–482 (1999).
Itô, Y. Behaviour and Social Evolution of Wasps (Oxford University Press, Oxford, 1993).
Packer, L. Multiple-foundress associations in sweat bees. In Queen Number and Sociality in Insects (ed. Keller, L.) 215–233 (Oxford University Press, Oxford, 1993).
Schwarz, M. P., Bull, N. J. & Hogendoorn, K. Evolution of sociality in the allodapine bees: A review of sex allocation, ecology and evolution. Insectes Soc.45, 349–368 (1998).
Shellman-Reeve, J. S. The spectrum of eusociality in termites. In The Evolution of Social Behavior in Insects and Arachnids (eds Choe, J. C. & Crespi, B. J.) 52–93 (Cambridge University Press, Cambridge, 1997).
Thorne, B. L. Evolution of eusociality in termites. Annu. Rev. Ecol. Evol. Syst.28, 27–54 (1997).
Hölldobler, B. & Wilson, E. O. The Ants (Springer, Berlin, 1990).
Schmid-Hempel, P. Parasites in Social Insects (Princeton University Press, Princeton, 1998).
Tschinkel, W. R. The Fire Ants (Harvard University Press, Cambridge, 2006).
Cole, B. J. The ecological setting of social evolution. In Organization of Insect Societies (eds Gadau, J. & Fewell, J.) 74–104 (Harvard University Press, Cambridge, 2009).
Johnson, R. A. Colony founding by pleometrosis in the semi-claustral seed-harvester ant Pogonomyrmex calfornicus (Hymenoptera: Formicidae). Anim. Behav.68, 1189–1200 (2004).
Tschinkel, W. R. An experimental study of pleometrotic colony founding in the fire ant, Solenopsis invicta: What is the basis for association?. Behav. Ecol. Sociobiol.43, 247–257 (1998).
Jerome, C. A., McInnes, D. A. & Adams, E. S. Group defense by colony-founding queens in the fire ant Solenopsis invicta. Behav. Ecol.9, 301–308 (1998).
Helms Cahan, S. & Julian, G. E. Fitness consequences of cooperative colony founding in the desert leaf-cutter ant Acromyrmex versicolor. Behav. Ecol.10, 585–591 (1999).
Adams, E. S. & Tschinkel, W. R. Effects of foundress number on brood raids and queen survival in the fire ant Solenopsis invicta. Behav. Ecol. Sociobiol.37, 233–242 (1995).
Clark, R. M. & Fewell, J. H. Social dynamics drive selection in cooperative associations of ant queens. Behav. Ecol.25, 117–123 (2014).
Offenberg, J., Peng, R. & Nielsen, M. Development rate and brood production in haplo- and pleometrotic colonies of Oecophylla smaragdina. Insectes Soc.59, 307–311 (2012).
Rissing, S. W. & Pollock, G. B. An experimental analysis of pleometric advantage in the desert seed-harvester ant Messor pergandei (Hymenoptera; Formicidae). Insectes Soc.38, 205–211 (1991).
Sasaki, K., Jibiki, E., Satoh, T. & Obara, Y. Queen phenotype and behaviour during cooperative colony founding in Polyrhachis moesta. Insectes Soc.52, 19–25 (2005).
Waloff, N. The effect of the number of queens of the ant Lasius flavus (Fab.) (Hym. Formicidae) on their survival and on the rate of development of the first brood. Insectes Soc.4, 391–408 (1957).
Bartz, S. H. & Hölldobler, B. Colony founding in Myrmecocystus mimicus Wheeler (Hymenoptera, Formicidae) and the evolution of foundress associations. Behav. Ecol. Sociobiol10, 137–147 (1982).
Helms Cahan, S. Ecological variation across a transition in colony-founding behavior in the ant Messor pergandei. Oecologia129, 629–635 (2001).
Sommer, K. & Hölldobler, B. Colony founding by queen association and determinants of reduction in queen number in the ant Lasius niger. Anim. Behav.50, 287–294 (1995).
Tschinkel, W. R. & Howard, D. F. Colony founding by pleometrosis in the fire ant, Solenopsis invicta. Behav. Ecol. Sociobiol12, 103–113 (1983).
Herbers, J. M. Nest site limitation and facultative polygyny in the ant Leptothorax longispinosus. Behav. Ecol. Sociobiol19, 115–122 (1986).
Nonacs, P. Queen condition and alate density affect pleometrosis in the ant Lasius pallitarsis. Insectes Soc.39, 3–13 (1992).
Masoni, A. et al. Pleometrotic colony foundation in the ant Crematogaster scutellaris (Hymenoptera: Formicidae): Better be alone than in bad company. Myrmecol. News25, 51–59 (2016).
Sommer, K. & Hölldobler, B. Pleometrosis in Lasius niger. In Biology and Evolution of Social Insects (ed. Billen, J.) 47–50 (Leuven University Press, Leuven, 1992).
Pfennig, D. W. Absence of joint nesting advantage in desert seed harvester ants: Evidence from a field experiment. Anim. Behav.49, 567–575 (1995).
Tschinkel, W. R. Brood raiding and the population dynamics of founding and incipient colonies of the fire ant Solenopsis invicta. Ecol. Entomol.17, 179–188 (1992).
Helms Cahan, S. & Fewell, J. H. Division of labor and the evolution of task sharing in queen associations of the harvester ant Pogonomyrmex californicus. Behav. Ecol. Sociobiol.56, 9–17 (2004).
Helmkampf, M., Mikheyev, A. S., Kang, Y., Fewell, J. & Gadau, J. Gene expression and variation in social aggression by queens of the harvester ant Pogonomyrmex californicus. Mol. Ecol.25, 3716–3730 (2016).
Overson, R. P., Gadau, J., Clark, R. M., Pratt, S. C. & Fewell, J. H. Behavioral transitions with the evolution of cooperative nest founding by harvester ant queens. Behav. Ecol. Sociobiol.68, 21–30 (2014).
Shaffer, Z. et al. The foundress’s dilemma: Group selection for cooperation among queens of the harvester ant, Pogonomyrmex californicus. Sci. Rep.6, 29828 (2016).
Aron, S., Steinhauer, N. & Fournier, D. Influence of queen phenotype, investment and maternity apportionment on the outcome of fights in cooperative foundations of the ant Lasius niger. Anim. Behav.77, 1067–1074 (2009).
Brütsch, T., Avril, A. & Chapuisat, M. No evidence for social immunity in co-founding queen associations. Sci. Rep.7, 16262 (2017).
Chérasse, S. & Aron, S. Measuring inotocin receptor gene expression in chronological order in ant queens. Horm. Behav.96, 116–121 (2017).
Dreier, S. & d’Ettorre, P. Social context predicts recognition systems in ant queens. J. Evol. Biol.22, 644–649 (2009).
Holman, L., Dreier, S. & d’Ettorre, P. Selfish strategies and honest signalling: Reproductive conflicts in ant queen associations. Proc. R. Soc. B277, 2007–2015 (2010).
Pull, C. D. & Cremer, S. Co-founding ant queens prevent disease by performing prophylactic undertaking behaviour. BMC Evol. Biol.17, 219 (2017).
Pull, C. D., Hughes, W. H. O. & Brown, M. J. F. Tolerating an infection: An indirect benefit of co-founding queen associations in the ant Lasius niger. Naturwissenschaften100, 1125–1136 (2013).
Bernasconi, G. & Keller, L. Phenotype and individual investment in cooperative foundress associations of the fire ant, Solenopsis invicta. Behav. Ecol9, 478–485 (1998).
Bernasconi, G. & Keller, L. Effect of queen phenotype and social environment on early queen mortality in incipient colonies of the fire ant, Solenopsis invicta. Anim. Behav.57, 371–377 (1999).
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