Giribet, G. & Edgecombe, G. D. Current understanding of Ecdysozoa and its internal phylogenetic relationships. Integr Comp Biol 57, 455–466 (2017).
Guidetti, R., Altiero, T. & Rebecchi, L. On dormancy strategies in tardigrades. J. Insect. Physiol. 57, 567–576 (2011).
Hashimoto, T. et al. Extremotolerant tardigrade genome and improved radiotolerance of human cultured cells by tardigrade-unique protein. Nat. Comm. 7, 12808 (2016).
Tsujimoto, M., Imura, S. & Kanda, H. Recovery and reproduction of Antarctic tardigrade retrieved from a moss sample frozen for over 30 years. Cryobiol. 72, 78–81 (2016).
Jönsson, K. I. Radiation tolerance in tardigrades: current knowledge and potential applications in medicine. Cancers 11, 1333 (2019).
Heidemann, N. W. T. et al. Osmotic stress tolerance in semi-terrestrial tardigrades. Zool. J. Linn. Soc. 178, 912–918 (2016).
Lewandowski, M. et al. Message in a stainless steel bottle thrown into deep geological time. Gondwana Res. 52, 139–141 (2017).
Convey, P. & McInnes, S. J. Exceptional tardigrade-dominated ecosystems in Ellsworth Land, Antarctica. Ecology 86, 519–527 (2005).
Adams, B. J. et al. Diversity and distribution of Victoria Land biota. Soil Biol. Biochem. 38, 3003–3018 (2006).
Sohlenius, B. & Boström, S. Species diversity and random distribution of microfauna in extremely isolated habitable patches on Antarctic nunataks. Polar Biol. 31, 817–825 (2008).
Guidetti, R., Rebecchi, L., Cesari, M. & McInnes, S. J. Mopsechiniscus franciscae, a new species of a rare genus of Tardigrada from continental Antarctica. Polar Biol. 37, 1221–1233 (2014).
Tsujimoto, M., Suzuki, A. C. & Imura, S. Life history of the Antarctic tardigrade, Acutuncus antarcticus, under a constant laboratory environment. Polar Biol. 38, 1575–1581 (2015).
Vecchi, M. et al. Integrative systematic studies on tardigrades from Antarctica identify new genera and new species within Macrobiotoidea and Echiniscoidea. Invertebr. Syst. 30, 303–322 (2016).
Velasco-Castrillón, A., Gibson, J. A. E. & Stevens, M. I. A review of current Antarctic limno-terrestrial microfauna. Polar Biol. 37, 1517–1531 (2014).
Guidetti, R., Massa, E., Bertolani, R., Rebecchi, L. & Cesari, M. Increasing knowledge of Antarctic biodiversity: new endimic taxa of tardigrades (Eutardigrada; Ramazzottiae) and their evolutionary relationships. Syst. Biodivers. 17, 573–593 (2019).
Kaczmarek, Ł. et al. New records of Antarctic Tardigrada with comments on interpopulation variability of the Paramacrobiotus fairbanksi Schill, Förster, Dandekar and Wolf, 2010. Diversity 12, 108 (2020).
Dastych, H. W. S. Tardigrada. Acta. Zool. Cracov. 28, 169–214 (1985).
Dastych, H. An annotated list of Alaskan Tardigrada. Pol. Polar Res. 3, 95–102 (1982).
Zawierucha, K., Coulson, S. J., Michalczyk, Ł. & Kaczmarek, Ł. Current knowledge of the Tardigrada of Svalbard with the first records of water bears from Nordaustlandet (High Arctic). Polar Res. 32, https://doi.org/10.3402/polar.v32i0.20886 (2013).
Trygvadóttir, B. V. & Kristensen, R. M. A zoogeographic study of the limnoterrestrial tardigrade fauna on the Faroe Islands. J. Limnol. 72(s1), 113–122 (2013).
Ochyra, R. & Vàňa, J. The hepatics of King George Island, South Shetlands, Antarctica, with particular reference to the Admiralty Bay region. Pol. Polar Res. 10, 183–210 (1989).
Pudełko, R. Two new topographic maps for sites of scientific interest on King George Island, West Antarctica. Pol. Polar Res. 29, 291–297 (2008).
Birkenmajer, K., Francalanci, L. & Peccerillo, A. Petrological and geochemical constraints on the genesis of Mesozoic-Cenozoic magmatism of King George Island, South Shetland Islands, Antarctica. Antarct. Sci. 3, 293–308 (1991).
Ferron, F. A., Simões, J. C., Aquino, F. E. & Setzer, A. W. Air temperature time series for King George Island, Antarctica. Pesqui. Antart. Bras. 4, 155–169 (2004).
Kejna, M., Araźny, A. & Sobota, I. Climatic change on King George Island in the years 1948-2011. Pol. Polar Res. 34, 213–235 (2013).
Kim, J. H., Ahn, I.-Y., Lee, K. S., Chung, H. & Choi, H.-G. Vegetation of Barton Peninsula in the neighbourhood of King Sejong Station (King George Island, maritime Antarctic). Polar Biol. 30, 903–916 (2007).
Bick, A. & Arlt, G. Description of intertidal macro- and meiobenthic assemblages in Maxwell Bay, King George Island, South Shetland Islands, Southern Ocean. Polar Biol. 36, 673–689 (2013).
Krzeminska, M., Sicinski, J. & Kuklinski, P. Biodiveristy and biogeographic affiliation of Bryozoa from King George Island (Antarctica). Syst. Biodivers. 16, 576–586 (2018).
Segadilha, J. L. & Lavrado, H. P. Tanaidacea fauna (Peracarida, Crustacea) from the shallow sublittoral zone of Admiralty Bay, King George Island, Antarctica, with new records. Polar Biol. 41, 589–597 (2018).
Iakovenko, N. S. et al. Antarctic bdelloid rotifers: diversity, endemism and evolution. Hydrobiologia 761, 5–43 (2015).
Mouratov, S., Lahav, I., Barness, G. & Steinberger, Y. Preliminary study of the soil nematode community at Machu Picchu Station, King George Island, Antarctica. Polar Biol. 24, 545–548 (2001).
Shimada, K., Pan, C. & Ohyama, Y. Variation in summer cold-hardiness of the Antarctic oribatid mite Alaskozetes antarcticus from contrasting habitats on King George Island. Polar Biol. 12, 701–706 (1992).
Janiec, K. The comparison of freshwater invertebrates of Spitsbergen (Arctic) and King George Island (Antarctic). Pol. Polar Res. 17, 173–202 (1996).
Dastych, H. The tardigrada from Antarctic with descriptions of several new species. Acta Zool. Cracov. 27, 377–436 (1984).
Kaczmarek, Ł. et al. Tardigrades from Larus dominicanus Lichtenstein, 1823 nests on the Argentine Islands (maritime Antarctic). Polar Biol. 41, 283–301 (2018).
Utsugi, K. & Ohyama, Y. Antarctic tardigrada III. Fildes Peninsula of King George Island. Proc. NIPR Symp. Polar. Biol. 6, 139–151 (1993).
Bertolani, R., Rebecchi, L. & Claxton, S. K. Phylogenetic significance of egg shell variation in tardigrades. Zool. J. Linnean Soc. 116, 139–148 (1996).
Murray, J. Scotish Tardigrada collected by the Lake Survey. Proc. R. Soc. Edinb. 45, 641–668 (1907).
Guidetti, R., Altiero, T. & Hansen, J. G. A new species of freshwater tardigrades from Disko Island (Greenland) increases an unsolved paradox in tardigrade systematics. Hydrobiologia 558, 69–79 (2006).
Dastych, H. Ramazzottius agannae sp. nov., a new tardigrade species from the nival zone of the Austrian Central Alps (Tardigrada). Entomol. Mitt. Zool. Mus. Hamburg 15, 237–253 (2011).
Hansen, J. G. & Katholm, A. K. A study of the genus Amphibolus from Disko Island with special attention on the life cycle of Amphibolus nebulosus (Eutardigrada: Eohypsibiidae). In: Hansen JG (ed) Arctic biology field course Quqertarsuaq, Copenhagen, Zoological Museum, University of Copenhagen, pp 129-163 (2002).
Stec, D., Morek, W. & Gąsiorek, P. Kaczmarek, Ł. & Michalczyk, Ł. Determinants and taxonomic consequences of extreme egg shell variability in Ramazzottius subanomalus (Biserov, 1985) (Tardigrada). Zootaxa 4208, 176–188 (2016).
Zawierucha, K. & Kolicka, M. & Kaczmarek, Ł. Re-description of the Arctic tardigrade Tenuibiotus voronkovi (Tumanov, 2007) (Eutardigrada; Macrobiotidea), with the first molecular data for the genus. Zootaxa 4196, 498–510 (2016).
Schuster, R. O., Nelson, D. R., Grigarick, A. A. & Christenberry, D. Systematic criteria of the Eutardigrada. T. Am. Microsc. Soc. 99, 284–303 (1980).
Perry, E. & Miller, W. R. & Kaczmarek, Ł. Recommended abbreviation for the names of genera of the phylum Tardigrada. Zootaxa 4608, 145–154 (2019).
McInnes, S. J. Zoogeographic distribution of terrestrial/freshwater tardigrades from current literature. J. Nat. Hist. 28, 257–352 (1994).
McInnes, S. J. Tardigrades from Signy Island, South Orkney Islands, with particular references to freshwater species. J. Nat. Hist. 29, 1419–1445 (1995).
Heywood, R. A limnological survey of the Ablation Point Area, Alexander Island, Antarctica. Philos. Trans. R. Soc. Lond. B Biol. Sci. 279, 39–54 (1977).
Argue, C. W. Tardigrades from New Brunswick, Canada. 2. Can. J. Zool. 50, 87–94 (1972).
Pilato, G. & Binda, M. G. Dactylobiotus caldarellai, nuova specie di eutardigrado della Terra del Fuoco. Animalia 21, 87–91 (1994).
Doyère, M. L. Mémoire sur les Tardigrades. Ann. Sci. Nat. Ser. 2 T. 14, 269–361 (1840).
Richters, F. Tardigrada in Handbuch der Zoologie (ed. Kükenthal, W. & Krumbach, T.) 1-68 (Walter de Gruyer & Co., 1926).
Marley, N. J., McInnes, S. J. & Sands, C. J. Phylum Tardigrada: reevaluation of the Parachela. Zootaxa 2819, 51–64 (2011).
Guidetti, R., Rebecchi, L. & Bertolani, R. Cuticle structure and systematics of the Macrobiotidae (Tardigrada, Eutardigrada). Acta Zool. 81, 27–36 (2000).
Schuster, R. O., Toftner, E. C. & Grigarick, A. A. Tardigrada of Pope Beach, Lake Tahoe, California. Wasmann J. Biol. 35, 115–136 (1977).
Gąsiorek, P., Stec, D. & Morek, W. & Michalczyk, Ł. Deceptive conservatism of claws: distinct phyletic lineages concealed within Isohypsibioidea (Eutardigrada) revealed by molecular and morphological evidence. Contrib. Zool. 88, 78–132 (2019).
Bertolani, R. 15. Tardigradi (Tardigrada) in Guide per il riconosciment o delle specie animali delle acque interne Italiane Consiglio Nazionale Delle Ricerche, (1982).
Guidetti, R., Gandolfi, A., Rossi, V. & Bertolani, R. Phylogenetic analysis of Macrobiotidae (Eutardigrada, Parachela): a combined morphological and molecular approach. Zool. Scr. 34, 235–244 (2005).
Bertolani, R. et al. Phylogeny of Eutardigrada: New molecular data and their morphological support lead to the identification of new evolutionary lineages. Mol. Phyl. Evol. 76, 110–126 (2014).
Cesari, M., Altiero, T. & Rebecchi, L. Identification of the trehalose-6-phosphate synthase (tps) gene in desiccation tolerant and intolerant tardigrades. Ital. J. Zool. 79, 530–540 (2012).
Jørgensen, A., Faurby, S., Hansen, J. G., Møbjerg, N. & Kristensen, R. M. Molecular phylogeny of Arthrotardigrada (Tardigrada). Mol. Phyl. Evol. 54, 1006–1015 (2010).
Chen, L. –J., Romano, F., Sewell, S. & Murdock, C. https://www.ncbi.nlm.nih.gov/nuccore/GQ925676.1/ (2009).
Guil, N., Jørgensen, A. & Kristensen, R. M. An upgraded comprehensive multilocus phylogeny of the Tardigrada tree of life. Zool Scr. 48, 120–137 (2019).
Sands, C. J., Convey, P., Linse, K. & McInnes, S. J. Assessing meiofaunal variation among individuals utilising morphological and molecular approaches: an example using the Tardigrada. BMC. Ecol. 8, 7, https://doi.org/10.1186/1472-6785-8-7 (2008).
Kaczmarek, Ł., Michalczyk, Ł. & Eggermont, H. Dactylobiotus luci, a new freshwater tardigrade (Eutardigrada: Macrobiotidae) from the Rwenzori Mountains, Uganda. Afr. Zool. 43, 150–155 (2008).
Kaczmarek, Ł., Schabetsberger, R., Litwin, M. & Michalczyk, Ł. A new freshwater eutardigrade from Fiji and Vanuatu (Oceania), with remarks on the genus Dactylobiotus. N. Zeal. J. Zool. 39, 311–318 (2012).
Biserov, V. The tardigrada of the Komandorskiye Islands, with a description of Dactylobiotus dervizi sp. n. (Eutardigrada, Macrobiotidae). Entomol. Mitt. Zool. Mus. Hamburg. 12, 327–336 (1998).
Beasley, C. W., Miller, W. R. & Shively, S. A new freshwater Tardigrada Dactylobiotus kansae sp. n. (Eutardigrada: Parachela: Murrayidae) from Kansas. USA Proc. Biol. Soc. Washington. 122, 460–463 (2009).
Thulin, G. Beiträge zur Kenntnis der Tardigradenfauna Schwedens. Ark. F. Zoologi. 7, 1–60 (1911).
Degma, P., Bertolani, R. & Guidetti, R. Actual checklist of Tardigrade species. http://www.tardigrada.modena.unimo.it/miscellanea/Actual%20checklist%20of%20Tardigrada.pdf, pp.46 (2018).
Dujardin, F. Obseravations zoologiques, II: Sure les Tardigrades et sur une espece a longs pieds vivant dans l’eau de mer. Ann. Sci. Nat. Zool. Ser.3 T. 15, 158–166 (1851).
Binda, M. G. & Pilato, G. Dactylobiotus lombardoi sp. n. (Eutardigrada: Macrobiotidae) from Tierra del Fuego, with a key to the Dactylobiotus-species. Zool. Anz. 238, 147–155 (1999).
Yang, T. Three new species and six new records of the class Eutardigrada (Tardigrada) from China (Parachela: Macrobiotidae, Hypsibiidae). Acta Zootax. Sin. 24, 444–453 (1999).
Yang, T. Three new species and one new record of the Tardigrada from China. Acta Hydrobiol. Sin. 26, 505–508 (2002).
McInnes, S. J., Pugh, P. J. A. & Robinson, K. Low temperature SEM: how eggs of a tardigrade respond to humidity changes. Microsc. Anal. 59, 37–39 (1997).
Mapalo, M. A., Stec, D. & Mirano-Bascos, D. & Michalczyk, Ł. Mesobiotus philippinicus sp. nov., the first limnoterrestrial tardigrade from the Philippines. Zootaxa 4126, 411–426 (2016).
Mapalo, M. A, Stec, D., Mirano-Bascos, D. & Michalczik, Ł. An integrative description of a limnoterrestrial tardigrade from the Philippines, Mesobiotus insanis, new species (Eutardigrada: Macrobiotidae: harmsworthi group). Raffles B. Zool. 65, 440–454 (2017).
Stec, D., Roszkowska, M., Kaczmarek, Ł. & Michalczyk, Ł. Paramacrobiotus lachowskae, a new species of Tardigrada from Colombia (Eutardigrada: Parachela: Macrobiotidae). New Zeal. J. Zool. https://doi.org/10.1080/03014223.2017.1354896 (2017).
Kaczmarek, Ł., Gawlak, M., Bartels, P. J., Nelson, D. R. & Roszkowska, M. Revision of the Genus Paramacrobiotus Guidetti et al., 2009 with the description of a new species, re-descriptions and a key. Ann. Zool. 67, 627–656 (2017).
Stec, D., Morek, W., Gąsiorek, P., Blagden, B. & Michalczyk, Ł. Description of Macrobiotus scoticus sp. nov. (Tardigrada: Macrobiotidae: hufelandi group) from Scotland by means of integrative taxonomy. Ann. Zool. 67, 181-197.
Stec, D. & Kristensen, R. M. An integrative description of Mesobiotus ethiopicus sp. nov. (Tardigrada: Eutardigrada: Parachela: Macrobiotidae: harmsworthi group) from the northern Afrotropic region. Turk. J. Zool. 41, 800–811 (2017).
Fontoura, P., Rubal, M. & Veiga, P. Two new species of Tardigrada (Eutardigrada: Ramazzottidae, Macrobiotidae) from the supralittoral zone of the Atlantic Iberian Peninsula rocky shores. Zootaxa 4263, 450–466 (2017).
Pennak, R. W. Fresh-water invertebrates of the United States. (Ronald Press, 1953).
Baumann, H. Lebenslauf und Lebensweise von Hypsibius (H.) oberhaeuseri Doyère (Tardigrada). Veröffentlichungen aus den Übersee-Museum Bermen 3, 245–258 (1966).
Schuetz, G. A one year study on the population dynamics of Milnesium tardigradum Doyère in the lichen Xanthoria parietina (L.) in Biology of tardigrades: Selected symposia and monographs (ed. Bertolani, R.) 217-228 (U.Z.I. Mucchi Editore, 1987).
Dastych, H. Two new Eutardigrada species from West Spitsbergen and the Tatra Mts. Bull. Soc. Amis. Sci. Lettr. Poznan 23, 195–200 (1983).
Ptatscheck, C., Gansfort, B. & Traunspurger, W. The extent of wind-mediated dispersal of small metazoans, focusing nematodes. Sci. Rep. 8, 6814, https://doi.org/10.1038/s41598-018-24247-8 (2018).
Zawierucha, K. et al. High mitochondrial diversity in a new water bear species (Tardigrada: Eutardigrada) from mountain glaciers in Central Asia, with the erection of a new genus. Cryoconicus. Ann. Zool. 68, 179–201 (2018).
Rivas, J. A. Jr., Schröder, T., Gill, T. E., Wallace, R. L. & Walsh, E. J. Anemochory of diapausing stages of microinvertebrates in North American drylands. Freshw. Biol. 64, 1303–1314 (2019).
Mogle, M. J., Kimball, S. A., Miller, W. R. & McKown, R. D. Evidence of avian-mediated long distance dispersal in American tardigrades. Peer J 6, e5035, https://doi.org/10.7717/peerj.5035 (2018).
Szymańska, B. Encystment in the tardigrade Dactylobiotus dispar (Murray 1907) (Tardigrada: Eutardigrada). Zool. Pol. 40, 91–102 (1995).
Poprawa, I., Hyra, M. & Rost-Roszkowska, M. M. Germ cell cluster organization and oogenesis in the tardigrade Dactylobiotus parthenogeneticus Bertolani, 1982 (Eutardigrada, Murrayidae). Protoplasma 252, 1019–1029 (2015).
Morek, D. et al. An experimental test of tardigrade preparation methods for light microscopy. Zool. J. Linn. Soc. 178, 785–793 (2016).
Shirayama, Y., Kaku, T. & Higgins, R. P. Double-sided microscopic observation of meiofauna using HS-slide. Benthos. Res. 44, 41–44 (1993).
Mitchell, C. & Miller, W. A simple SEM (Scanning Electron Microscope) preparation protocol for tardigrades. J. Pa. Acad. Sci. 81, 86–90 (2008).
Pilato, G. Analisi di nuovi caratteri nello studio degli Eutardigradi. Animalia 8, 51–57 (1981).
Bartels, P. J., Nelson, D. R. & Exline, R. P. Allometry and the removal of body size effects in the morphometric analysis of tardigrades. J. Zool. Syst. Evol. Res. 49, 17–25 (2011).
Rammazzotti, G. & Maucci, W. Il Phylum Tardigrada. Terza edizione riveduta e aggiornata English translated by Beasley CW. Mem. Ist. Ital. Idrobiol. Dott. Marco De Marchi 41 (1983).
Kaczmarek, Ł. & Michalczyk, Ł. The Macrobiotus hufelandi group (Tardigrada) revisited. Zootaxa 4363, 101–123 (2017).
Pilato, G. & Binda, M. G. Definition of families, subfamilies, genera and subgenera of the Eutardigrada, and keys to their identification. Zootaxa 2404, 1–54 (2010).
Hall, T. A. BioEdit: a user-friendly biological sequence alignment editor and analysis program for Windows 95/98/NT. Nucleic Acids Symp. Ser. 41, 95–98 (1999).
Kunmar, S., Stecher, G., Li, M., Knyaz, C. & Tamura, K. MEGA X: Molecular Evolutionary Genetics Analysis across computing platforms. Mol. Biol. Evol. 35, 1547–1549 (2018).
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