No protofeathers on pterosaurs

Wellnhofer, P. The Illustrated Encyclopedia of Pterosaurs (Salamander Books, 1991).

Unwin, D. M. The Pterosaurs from Deep Time (Pi, 2005).

Kellner, A. W. A. et al. The soft tissue of Jeholopterus (Pterosauria, Anurognathidae, Batrachognathinae) and the structure of the pterosaur wing membrane. Proc. R. Soc. B 277, 321–329 (2010).

Article  Google Scholar 

Witton, M. P. Pterosaurs. Natural History, Evolution, Anatomy (Princeton Univ. Press, 2013).

Yang, Z. et al. Pterosaur integumentary structures with complex feather-like branching. Nat. Ecol. Evol. 4, 24–30 (2019).

Article  Google Scholar 

Xu, X. et al. An integrative approach to understanding bird origins. Science 346, 1253293 (2014).

Article  Google Scholar 

Benton, M. J., Dhouially, D., Jiang, B. & McNamara, M. The early origin of feathers. Trends Ecol. Evol. 34, 856–859 (2019).

Article  Google Scholar 

Xu, X. Feather evolution: Looking up close and through deep time. Sci. Bull. 64, 563–564 (2019).

Article  Google Scholar 

Unwin, D. M. & Bakhurina, N. N. Sordes pilosus and the nature of the pterosaur flight apparatus. Nature 371, 62–64 (1994).

Article  Google Scholar 

Frey, E., Tischlinger, H., Buchy, M. C. & Martill, D. M. in Evolution and Palaeobiology of Pterosaurs Vol. 217 (eds Buffetaut, E. & Mazin, J.-M.) 233–266 (Geological Society of London, Special Publications, 2003).

Li, Q. et al. Melanosome evolution indicates a key physiological shift within feathered dinosaurs. Nature 507, 350–353 (2014).

CAS  Article  Google Scholar 

Pinheiro, F. L. et al. Chemical characterization of pterosaur melanin challenges color inferences in extinct animals. Sci. Rep. 9, 15947 (2019).

Article  Google Scholar 

Campione, N. E., Barrett, P. M. & Evans, D. C. in The Evolution of Feathers (eds Foth, C. & Rauhut, O. W. M.) 213–243 (Springer, 2020).

Czerkas, S. A. & Ji, Q. in Feathered Dinosaurs and the Origin of Flight (ed. Czerkas, S. J.) 15–41 (The Dinosaur Museum, 2002).

Ji, Q. & Yuan, C. Discovery of two kinds of protofeathered pterosaurs in the Mesozoic Daohugou biota in the Ningcheng region and its stratigraphic and biologic significances. Geol. Rev. 48, 221–224 (2002).

Google Scholar 

Wang, X. L., Zhou, Z. H., Zhang, F. C. & Xu, X. A nearly completely articulated rhamphorhynchoid pterosaur with exceptionally well-preserved wing membranes and ‘hairs’ from Inner Mongolia, northeast China. Chin. Sci. Bull. 47, 226–230 (2002).

Article  Google Scholar 

Jäger, K. R. K., Tischlinger, H., Oleschinski, G. & Sander, P. M. Goldfuß was right: soft part preservation in the Late Jurassic pterosaur Scaphognathus crassirostris revealed by reflectance transformation imaging and ultraviolet light and the auspicious beginnings of paleo-art. Palaeontol. Electron. 21.3.3T (2019).

Lü, J. & Hone, D. W. E. A new Chinese anurognathid pterosaur and the evolution of pterosaurian tail lengths. Acta Geol. Sin. 86, 1317–1325 (2012).

Article  Google Scholar 

Bennett, S. C. New interpretation of the wings of the pterosaur Rhamphorhynchus muensteri based on the Zittel and Marsh specimens. J. Paleontol. 89, 845–869 (2016).

Article  Google Scholar 

Hone, D., Henderson, D. M., Therrien, F. & Habib, M. B. A specimen of Rhamphorhynchus with soft tissue preservation, stomach contents and a putative coprolite. PeerJ 3, e1191 (2015).

Article  Google Scholar 

Pan, Y. et al. Molecular evidence of keratin and melanosomes in feathers of the Early Cretaceous bird Eoconfuciusornis. Proc. Natl Acad. Sci. USA 113, E7900–E7907 (2016).

CAS  Article  Google Scholar 

Alibardi, L. Adaptation to the land: the skin of reptiles in comparison to that of amphibians and endotherm amniotes. J. Exp. Zool. 298B, 12–41 (2003).

CAS  Article  Google Scholar