Wellnhofer, P. The Illustrated Encyclopedia of Pterosaurs (Crescent Books, 1991).
Unwin, D. M. The pterosaurs from deep time (Pi Press, 2005).
Witton, M. P. Pterosaurs: Natural History (Anatomy (Princeton University Press, 2013).
Google Scholar
Williams, C. J. et al. Helically arranged cross struts in azhdarchid pterosaur cervical vertebrae and their biomechanical implications. iScience 24, 102338 (2021).
Google Scholar
Bestwick, J., Unwin, D. M., Butler, R. J. & Purnell, M. A. Dietary diversity and evolution of the earliest flying vertebrates revealed by dental microwear texture analysis. Nat. Commun. 11, 5293 (2020).
Google Scholar
Ryang, W. H. Characteristics of strike-slip basin formation and sedimentary fills and the Cretaceous small basins of the Korean Peninsula. J. Geo. Soc. Korea 49, 31–45 (2013).
Google Scholar
Kim, B. G. & Park, B. G. Geological report of the Dongbok sheet (1:50,000) (Geological Survey of Korea, Seoul, 1966).
Lee, H., Sim, M. S. & Choi, T. Stratigraphic evolution of the northern part of the Cretaceous Neungju basin South Korea. Geosci. J. 23, 849–865 (2019).
Google Scholar
Paik, I. S., Huh, M., So, Y. H., Lee, J. E. & Kim, H. J. Traces of evaporites in Upper Cretaceous lacustrine deposits of Korea: Origin and paleoenvironmental implications. J. Asian Earth Sci. 30, 93–107 (2007).
Google Scholar
Cohen, K. M., Finney, S. M., Gibbard, P. L. & Fan, J.-X. The ICS international Chronostratigraphic chart. Episodes 36, 199–204 (2013).
Google Scholar
Calvo, J. O. & Lockley, M. G. The first pterosaur tracks from Gondwana. Cretac. Res. 22, 585–590 (2001).
Google Scholar
Kukihara, R. & Lockley, M. G. Fossil footprints from the dakota group (Cretaceous) john martin reservoir, bent county, Colorado: New insights into the paleoecology of the Dinosaur freeway. Cretac. Res. 33, 165–182 (2012).
Google Scholar
Lockley, M. & Schumacher, B. A new pterosaur swim tracks locality from the Cretaceous Dakota Group of eastern Colorado: implications for pterosaur swim track behavior. Fossil Footprints of Western North America. Bull. NM Mus. Nat. Hist. Sci, 365–371 (2014).
Smith, R. E., Martill, D. M., Unwin, D. M. & Steel, L. Edentulous pterosaurs from the Cambridge Greensand (Cretaceous) of eastern England with a review of Ornithostoma Seeley, 1871. Proc. Geol. Assoc. (2020).
Ibrahim, N., Unwin, D. M., Martill, D. M., Baidder, L. & Zouhri, S. A new pterosaur (Pterodactyloidea: Azhdarchidae) from the Upper Cretaceous of Morocco. PLoS ONE 5, e10875 (2010).
Google Scholar
Martill, D. M. & Ibrahim, N. An unusual modification of the jaws in cf. Alanqa, a mid-Cretaceous azhdarchid pterosaur from the Kem Kem beds of Morocco. Cretac. Res. 53, 59–67 (2015).
Google Scholar
Jacobs, M. L., Martill, D. M., Ibrahim, N. & Longrich, N. A new species of Coloborhynchus (Pterosauria, Ornithocheiridae) from the mid-Cretaceous of North Africa. Cretac. Res. 95, 77–88 (2019).
Google Scholar
Jacobs, M. L. et al. New toothed pterosaurs (Pterosauria: Ornithocheiridae) from the middle Cretaceous Kem Kem beds of Morocco and implications for pterosaur palaeobiogeography and diversity. Cretac. Res. 110, 104413 (2020).
Google Scholar
McPhee, J. et al. A new ? Chaoyangopterid (Pterosauria: Pterodactyloidea) from the Cretaceous Kem Kem beds of southern Morocco. Cretac. Res. 110, 104410 (2020).
Google Scholar
Martill, D. M. et al. A new tapejarid (Pterosauria, Azhdarchoidea) from the mid-Cretaceous Kem Kem beds of Takmout, southern Morocco. Cretac. Res. 112, 104424 (2020).
Google Scholar
Martill, D. M., Unwin, D. M., Ibrahim, N. & Longrich, N. A new edentulous pterosaur from the Cretaceous Kem Kem beds of south eastern Morocco. Cretac. Res. 84, 1–12 (2018).
Google Scholar
Smith, R. E. et al. Small, immature pterosaurs from the Cretaceous of Africa: implications for taphonomic bias and palaeocommunity structure in flying reptiles. Cretac. Res. 130, 105061 (2022).
Google Scholar
Smith, R. E., Martill, D. M., Kao, A., Zouhri, S. & Longrich, N. A long-billed, possible probe-feeding pterosaur (Pterodactyloidea: ?Azhdarchoidea) from the mid-Cretaceous of Morocco North Africa. Cretac. Res. 118, 104643 (2021).
Google Scholar
Kellner, A. W. A. et al. First complete pterosaur from the Afro-Arabian continent: insight into pterodactyloid diversity. Sci. Rep. 9, 17875 (2019).
Google Scholar
Elgin, R. A. & Frey, E. A new azhdarchoid pterosaur from the Cenomanian (Late Cretaceous) of Lebanon. Swiss J. Geosci. 104, 21–33 (2011).
Google Scholar
Averianov, A. O., Kurochkin, E. N., Pervushov, E. M. & Ivanov, A. V. Two bone fragments of ornithocheiroid pterosaurs from the Cenomanian of Volgograd Region, southern Russia. Acta Palaeontol. Pol. 50 (2005).
Averianov, A. & Kurochkin, E. A new pterosaurian record from the Cenomanian of the Volga region. Paleontol. J. 44, 695–697 (2010).
Google Scholar
Nessov, L. Flying reptiles from the Jurassic and cretaceous of the USSR and significance of their remains for the reconstruction of paleogeographical conditions. Vestn. Leningr. Gos. Univ. Ser. 7, 28 (1990).
Bakhurina, N. N. & Unwin, D. M. A survey of pterosaurs from the Jurassic and Cretaceous of the former Soviet Union and Mongolia. (1995).
Averianov, A. O. New records of azhdarchids (Pterosauria, Azhdarchidae) from the Late Cretaceous of Russia, Kazakhstan, and Central Asia. Paleontol. J. 41, 189–197 (2007).
Google Scholar
Averianov, A. Mid-Cretaceous ornithocheirids (Pterosauria, Ornithocheiridae) from Russia and Uzbekistan. Paleontol. J. 41, 79–86 (2007).
Google Scholar
Huh, M., Paik, I. S., Chung, C. H., Hwang, K. G. & Kim, B. S. Theropod tracks from Seoyuri in Hwasun, Jeollanamdo, Korea: occurrence and paleontological significance. J. Geo. Soc. Korea 39, 461–478 (2003).
Google Scholar
Huh, M. et al. Well-preserved theropod tracks from the Upper Cretaceous of Hwasun County, southwestern South Korea, and their paleobiological implications. Cretac. Res. 27, 123–138 (2006).
Google Scholar
Lockley, M. G., Huh, M. & Kim, B. S. Ornithopodichnus and pes-only sauropod Trackways from the Hwasun tracksite Cretaceous of Korea. Ichnos 19, 93–100 (2012).
Google Scholar
Hwang, K. G., Huh, M. & Paik, I. S. A unique trackway of small theropod from Seoyu-ri, Hwasun-gun Jeollanam province. J. Geo. Soc. Korea 42, 69–78 (2006).
Google Scholar
Kim, B. S. & Huh, M. Analysis of the acceleration phase of a theropod dinosaur based on a Cretaceous trackway from Korea. Palaeogeogr. Palaeoclimatol. Palaeoecol. 293, 1–8 (2010).
Google Scholar
Marchetti, L. et al. Defining the morphological quality of fossil footprints. Problems and principles of preservation in tetrapod ichnology with examples from the Palaeozoic to the present. Earth-Sci. Rev. 193, 109–145 (2019).
Google Scholar
Rodríguez-de La Rosa, R. A. Pterosaur tracks from the latest Campanian Cerro del Pueblo formation of southeastern Coahuila. Mexico. Geol. Soc. Spec. Publ. 271, 275–282 (2003).
Google Scholar
Lockley, M. G. & Meyer, C. Crocodylomorph trackways from the Jurassic to early cretaceous of North America and Europe: Implications for Ichnotaxonomy. Ichnos 11, 167–178 (2004).
Google Scholar
Ambroggi, R. & De Lapparent, A. Les empreintes de pas fossiles du Maestrichtien d’Agadir. Notes du Service Géologique du Maroc 10, 43–57 (1954).
Stokes, W. L. Pterodactyl tracks from the Morrison Formation. J. Paleontol. 31, 952–954 (1957).
Delair, J. Note on Purbeck fossil footprints, with descriptions of two hitherto unknown forms from Dorset. Proceedings of the Dorset Natural History and Archaeological Society. 92–100 (1963).
Hwang, K.-G., Huh, M. I. N., Lockley, M. G., Unwin, D. M. & Wright, J. L. New pterosaur tracks (Pteraichnidae) from the Late Cretaceous Uhangri Formation, southwestern Korea. Geol. Mag. 139, 421–435 (2002).
Google Scholar
Mazin, J.-M. & Pouech, J. The first non-pterodactyloid pterosaurian trackways and the terrestrial ability of non-pterodactyloid pterosaurs. Geobios 58, 39–53 (2020).
Google Scholar
Masrour, M., de Ducla, M., Billon-Bruyat, J.-P. & Mazin, J.-M. Rediscovery of the Tagragra tracksite (Maastrichtian, Agadir, Morocco): Agadirichnus elegans Ambroggi and Lapparent 1954 is Pterosaurian Ichnotaxon. Ichnos 25, 285–294 (2018).
Google Scholar
Wright, J. L., Unwin, D. M., Lockley, M. G. & Rainforth, E. C. Pterosaur tracks from the Purbeck limestone formation of Dorset England. Proc. Geol. Assoc. 108, 39–48 (1997).
Google Scholar
Lockley, M. G. et al. The fossil trackway Pteraichnusis pterosaurian, not crocodilian: Implications for the global distribution of pterosaur tracks. Ichnos 4, 7–20 (1995).
Google Scholar
Billon-Bruyat, J.-P. & Mazin, J.-M. The systematic problem of tetrapod ichnotaxa: the case study of Pteraichnus Stokes, 1957 (Pterosauria, Pterodactyloidae). Geol. Soc. Spec. Publ. 217, 315–324 (2003).
Google Scholar
Pascual Arribas, C. & Sanz Pérez, E. Huellas de Pterosaurios en el grupo Oncala (Soria, España). Pteraichnus palaciei-saenzi, nov. icnosp. Estudios Geol. 56, 73–100 (2000).
Calvo, M. M., Vidarte, C. F., Fuentes, F. M. & Fuentes, M. M. Huellas de Pterosaurios en la Sierra de Oncala (Soria, España). Nuevas icnoespecies: pteraichnus vetustior, Pteraichnus parvus. Pteraichnus manueli. Celtiberia 54, 471–490 (2004).
Fuentes Vidarte, C., Meijide Calvo, M., Meijide Fuentes, F. & Meijide Fuentes, M. Pteraichnus longipodus nov. icnosp. en la Sierra de Oncala (Soria, España). Studia Geologica Salmanticensia, 103–114 (2004).
Peng, B.-X., Du, Y.-S., Li, D.-Q. & Bai, Z.-C. The first discovery of the early Cretaceous Pterosaur track and its significance in Yanguoxia, Yongjing County, Gansu Province. Earth Sci.-J. China Univ. Geosci. 29, 21–24 (2004).
Lee, Y.-N., Lee, H.-J., Lü, J. & Kobayashi, Y. New pterosaur tracks from the Hasandong formation (Lower Cretaceous) of Hadong County South Korea. Cretac. Res. 29, 345–353 (2008).
Google Scholar
Lee, Y.-N., Azuma, Y., Lee, H.-J., Shibata, M. & Lü, J. The first pterosaur trackways from Japan. Cretac. Res. 31, 263–273 (2010).
Google Scholar
Chen, R. et al. Pterosaur tracks from the early late cretaceous of Dongyang City, Zhejiang Province China. Geol. Bull. China. 32, 693–698 (2013).
Google Scholar
Li, Y., Wang, X. & Jiang, S. A new pterosaur tracksite from the Lower Cretaceous of Wuerho, Junggar Basin, China: inferring the first putative pterosaur trackmaker. PeerJ 9, e11361 (2021).
Google Scholar
Ha, S. et al. Diminutive pterosaur tracks and trackways (Pteraichnus gracilis ichnosp. Nov.) from the lower Cretaceous Jinju formation, Gyeongsang basin. Korea. Cretac. Res. 131, 105080 (2021).
Google Scholar
Sánchez-Hernández, B., Przewieslik, A. G. & Benton, M. J. A reassessment of the Pteraichnus ichnospecies from the early Cretaceous of Soria Province Spain. J. Vertebr. Paleontol. 29, 487–497 (2009).
Google Scholar
Zhou, X. et al. A new darwinopteran pterosaur reveals arborealism and an opposed thumb. Curr. Biol. 31, 2429-2436.e2427 (2021).
Google Scholar
Lü, J. et al. Dragons of the Skies (recent advances on the study of pterosaurs from China) (Zhejiang Science and Technology Press, 2013).
Beccari, V. et al. Osteology of an exceptionally well-preserved tapejarid skeleton from Brazil: Revealing the anatomy of a curious pterodactyloid clade. PLoS ONE 16, e0254789 (2021).
Google Scholar
Lü, J. A new boreopterid pterodactyloid pterosaur from the Early Cretaceous Yixian Formation of Liaoning Province, northeastern China. Acta Geologica Sinica-English Edition 84, 241–246 (2010).
Google Scholar
Bennett, S. C. Terrestrial locomotion of pterosaurs: A reconstruction based on Pteraichnus trackways. J. Vertebr. Paleontol. 17, 104–113 (2010).
Google Scholar
Wang, X. & Lü, J. Discovery of a pterodactylid pterosaur from the Yixian Formation of western Liaoning China. Chin. Sci. Bull. 46, A3–A8 (2001).
Google Scholar
Frey, E. et al. A new specimen of nyctosaurid pterosaur, cf. Muzquizopteryx sp. from the Late Cretaceous of northeast Mexico. Revista mexicana de ciencias geológicas 29, 131–139 (2012).
Wu, W.-H., Zhou, C.-F. & Andres, B. The toothless pterosaur Jidapterus edentus (Pterodactyloidea: Azhdarchoidea) from the Early Cretaceous Jehol Biota and its paleoecological implications. PLoS ONE 12, e0185486 (2017).
Google Scholar
Lü, J. et al. The toothless pterosaurs from China. Acta Geol. Sin. 90, 2513–2525 (2016).
Zhang, X., Jiang, S., Cheng, X. & Wang, X. New Material of Sinopterus (Pterosauria, Tapejaridae) from the Early Cretaceous Jehol Biota of China. An. Acad. Bras. Cienc. 91 (2019).
Bestwick, J., Unwin, D. M., Butler, R. J., Henderson, D. M. & Purnell, M. A. Pterosaur dietary hypotheses: A review of ideas and approaches. Biol. Rev. 93, 2021–2048 (2018).
Google Scholar
Chen, H. et al. New anatomical information on Dsungaripterus weii Young, 1964 with focus on the palatal region. PeerJ 8, e8741 (2020).
Google Scholar
Li, D. et al. A manus dominated pterosaur track assemblage from Gansu, China: Implications for behavior. Sci. Bull. 60, 264–272 (2015).
Google Scholar
Masrour, M., Pascual-Arribas, C., de Ducla, M., Hernández-Medrano, N. & Pérez-Lorente, F. Anza palaeoichnological site. Late Cretaceous. Morocco. Part I. The first African pterosaur trackway (manus only). J. African Earth Sci. 134, 766–775 (2017).
Google Scholar
Bramwell, C. D. & Whitfield, G. R. Biomechanics of Pteranodon. Phil. Trans. R. Soc. Lond. B. 267, 503–581 (1974).
Google Scholar
Bennett, S. C. Terrestrial locomotion of pterosaurs: a reconstruction based on Pteraichnus trackways. J. Vertebr. Paleontol. 17, 104–113 (1997).
Google Scholar
Mazin, J.-M., Billon-Bruyat, J.-P., Hantzpergue, P. & Lafaurie, G. Ichnological evidence for quadrupedal locomotion in pterodactyloid pterosaurs: Trackways from the Late Jurassic of Crayssac (southwestern France). Geol. Soc. Spec. Publ. 217, 283–296 (2003).
Google Scholar
Henderson, D. M. Pterosaur body mass estimates from three-dimensional mathematical slicing. J. Vertebr. Paleontol. 30, 768–785 (2010).
Google Scholar
Lockley, M. G. & Wright, J. L. Pterosaur swim tracks and other ichnological evidnce of behaviour and ecology. Geol. Soc. Spec. Publ. 217, 297–313 (2003).
Google Scholar
Lockley, M., Mitchell, L. & Odier, G. P. Small Theropod track assemblages from middle Jurassic Eolianites of eastern Utah: Paleoecological insights from dune Ichnofacies in a transgressive sequence. Ichnos 14, 131–142 (2007).
Google Scholar
Fiorillo, A. R., Hasiotis, S. T., Kobayashi, Y. & Tomsich, C. S. A pterosaur manus track from Denali National park, Alaska Range, Alaska United States. Palaios 24, 466–472 (2009).
Google Scholar
Bell, P. R., Fanti, F. & Sissons, R. A possible pterosaur manus track from the late Cretaceous of Alberta. Lethaia 46, 274–279 (2013).
Google Scholar
Stinnesbeck, W. et al. Theropod, avian, pterosaur, and arthropod tracks from the uppermost Cretaceous Las Encinas Formation, Coahuila, northeastern Mexico, and their significance for the end-Cretaceous mass extinction. Geol. Soc. Am. Bull. 129, 331–348 (2017).
Google Scholar
Xing, L. et al. Late Cretaceous ornithopod-dominated, theropod, and pterosaur track assemblages from the Nanxiong Basin, China: New discoveries, ichnotaxonomy, and paleoecology. Palaeogeogr. Palaeoclimatol. Palaeoecol. 466, 303–313 (2017).
Google Scholar
Lockley, M. G., Gierlinski, G. D., Adach, L., Schumacher, B. & Cart, K. Newly discovered tetrapod ichnotaxa from the Upper Blackhawk Formation Utah. Bull. N. M. M. Nat. Hist. Sci. 79, 469–480 (2018).
Lockley, M. G. & Gillette, D. Pterosaur and bird tracks from a new Late Cretaceous locality in Utah. Verteb. Paleontol. Utah 99, 355–359 (1999).
Bennett, S. C. The ontogeny of Pteranodon and other pterosaurs. Paleobiology 19, 92–106 (1993).
Google Scholar
Bennett, S. C. Year-classes of pterosaurs from the Solnhofen Limestone of Germany: taxonomic and systematic implications. J. Vertebr. Paleontol. 16, 432–444 (1996).
Google Scholar
Chiappe, L. M., Codorniú, L., Grellet-Tinner, G. & Rivarola, D. Argentinian unhatched pterosaur fossil. Nature 432, 571–572 (2004).
Google Scholar
Codorniú, L., Chiappe, L. & Rivarola, D. Neonate morphology and development in pterosaurs: evidence from a Ctenochasmatid embryo from the Early Cretaceous of Argentina. Geol. Soc. Spec. Publ. 455, 83–94 (2018).
Google Scholar
Mickelson, D. L., Lockley, M. G., Bishop, J. & Kirkland, J. A New Pterosaur Tracksite from the Jurassic Summerville Formation, near Ferron Utah. Ichnos 11, 125–142 (2004).
Google Scholar
Source: Ecology - nature.com
