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    Device for automatic measurement of light pollution of the night sky

    For several years, systematic research has been carried out on the pollution of the night sky by artificial light in the city of Toruń11,36,38. The main objective is to monitor this phenomenon, including its spatial and temporal variability and the most important factors affecting it. Based on previous experience, an in-house measurement device was constructed to automate the process of data acquisition.Genesis of the projectThe first measurements pertaining to the phenomenon of night sky pollution in Toruń were made in autumn 2017, followed by regular observations using a handheld SQM photometer (Unihedron, Canada) as part of a project implemented in 2017–2018. To this end, a permanent measurement network distributed throughout the city was established, consisting of 24 locations. During a one night measurement session taking place during the astronomical night (there is no such period at the latitude of Toruń in the summer), sky brightness was measured at all sites. The results of spot monitoring were plotted using interpolation methods and visualisation tools available in GIS systems11,39, which helped to determine the spatial distribution and extent of night sky light pollution. The intensity of this phenomenon at each of the surveyed points was also explored in relation to the distinguished landcover categories and types of urban development.Repeatable measurements performed regularly over such a long period of time were characterised by significant limitations. One measurement session was very time-consuming, as it lasted about two hours, during which time all measurement locations were visited, covering a distance of almost 50 km each night by car. Despite the observance of all time frames and sticking to the plan of fieldwork, measurements were not carried out simultaneously at all the locations, which affected the results, especially during the night with changing cloud cover. Although the measurements were carried out with great consistency and care, they were performed in a spatial buffer of about 5 m, which could unintentionally slightly affect the obtained results. An additional limitation was also a one-time night measurement at one point, instead of a whole series of measurements at specific time intervals. Inaccuracies in the readings within a single session could have been caused by sudden changes in meteorological parameters. In the adopted procedure, it was not possible to carry out simultaneous measurements in identical time and weather conditions at all the locations, not to mention the involvement of the personnel in each tour of the measurement network points.Using the experience gained and after an analysis of the identified constraints and the technical capabilities at hand, work began in 2019 on developing a network for automatic remote monitoring of light pollution of the night sky in Toruń, based on designed in-house recording devices.Design, functional and utility features of the deviceTo enhance the research on light pollution in urban space, work has begun on the construction of a device that would perform automatic measurements, would be mobile, battery-powered and use long-range wireless communication. All the aforementioned features are in line with the strategy of Industry 4.0 and modern solutions proposed as part of the Smart City concept.The concept of Industry 4.0 assumes the more and more common use of process automation as well as the processing and exchange of data with the use of new transmission technologies26. LoRaWAN is one of the solutions used for communication of Internet of Things (IoT) devices, which supports the development of Smart Cities in the Smart Environment area. As a result, the interactivity, frequency, and scope of measurements carried out in urbanized areas are increased40,41.According to the developed project, the device was to serve as a meter of very low intensity light observed in the night sky. In this respect, it was necessary to use a sensor with technical parameters suitable for very accurate measurements of light intensity. To locally verify the weather conditions occurring during the operation of the device, it was decided to carry out additional simultaneous measurements of other environmental parameters—temperature and moisture content. The analysis of the spatial coverage of the study area indicated that 36 measurement devices should be deployed to provide full coverage of Toruń. The concept of creating an urban measurement network assumes the selection of points covering the whole city relatively evenly and representing different types of housing development and elements of land cover. It was assumed that measurements will be made only at night, between 21 p.m. and 6 a.m. on the following day, at 15 min intervals, and in addition, weather conditions will be recorded twice a day.Construction and technical parameters of the device, and selected characteristics of its componentsA prototype device meeting all the predefined functionalities was constructed based on available electronic modules. The B-L072Z-LRWAN Discovery developmental board from STMicroelectronics42 was selected as the main electronic component providing wireless communication. This board has an integrated LoRa communication module, enabling low-power wireless messaging, and also allows the board to enter a low-power state during hibernation, and thus target long-term battery-powered operation. This module is fully programmable, which enables future expansion of the set with other functionalities. The TSL2591 light sensor from AMS, which has high sensitivity and registration accuracy, was selected as a component implementing the light intensity measurement. Its great advantage is a wide measurement range of 188 μlx to 88 000 lx, sensitivity reaching 0.000377 lx, and a wide dynamic range (WDR) of 600 M:143. The sensor used has two diodes with different spectral properties. One of them registers visible light together with infrared (in the range from 400 to 1 100 nm), while the other is responsible for the registration of infrared light (between 500 and 1 100 nm). Thanks to this solution, we can use the results in various ways. The use of the formula provided by the manufacturer allows us to obtain spectral characteristics similar to the human eye. The presence of a compensating diode makes a difference compared to the sensor used in the SQM device, so the results obtained in the measurements may be slightly different.To measure additional environmental parameters, the X-NUCLEO-IKS01A2 development board from STMicroelectronics was used, which is connected to the STM32 microcontroller via the I2C interface44. This board enables the recording of a number of parameters, however, in the constructed device it is only responsible for reading the temperature and humidity of the environment. This results from the necessity to limit the size of the message packets sent, while at the same time improving the operating range and reducing the power consumption of the device.Once all the components had been selected, tested and integrated, the process of final connection and programming was carried out. The base of the device, i.e. the B-L072Z-LRWAN development board was connected to the X-NUCLEO-IKS01A2 environmental sensor board, using Arduino connectors. Using standard wires, a TSL2591 light sensor was added by connecting the corresponding I2C (SCL and SDA), power supply (VIN), sensor ground (GND) pins and the X-NUCLEO-IKS01A2 board.All components used were placed in a standard external casing with dimensions of 8.0 × 5.4 × 15.8 cm. In its lower part an opening was made for an external antenna, while in the upper part a specifically selected opening was cut out, protected with a glass pane, through which measurements are performed by the light sensor (Fig. 2).Figure 2(photo by Dominika Karpińska).Constructed device viewFull size imageFollowing the above steps, an automatic device was constructed to record light intensity in the lower troposphere, i.e. to measure the pollution of the night sky by artificial light coming from the Earth’s surface. Selected technical parameters of the device are presented in Table 1.Table 1 Selected technical parameters of the device for measuring light pollution of the night sky.Full size tableFlowchart of the system operationAfter constructing the device and writing the control software, the construction of the entire measurement system was started. Each of the measuring instruments is ultimately connected to the communication gateway using LoRa technology. A MultiTech communication gateway with a LoRaWAN module was used as an access device. To successfully connect the gateway to the measuring device, it was necessary to configure the communication gateway software. To this end, the information about the unique device number (Dev EUI) and the application key and its number (App EUI and App Key) was used. Once the unit is configured, it is possible to send data to the communication gateway and read them using NodeRED, a programming tool where data are redirected to a selected server, which stores all measurement results. Figure 3 shows a schematic representation of the constructed measurement system.Figure 3Schematic diagram of the measurement system.Full size image More

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    Soil meso- and micro-fauna community in response to bamboo-fungus agroforestry management

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    Study of cattle microbiota in different regions of Kazakhstan using 16S metabarcoding analysis

    Comparative characteristics of rations for feeding cattle from different regions of the Republic of Kazakhstan and the impact of animal feeding types on the faecal microbiotaDue to the huge differences in the natural and climatic conditions of Kazakhstan, animals from different regions of Kazakhstan were enrolled for this study. The difference in soil and climatic conditions of different zones has a significant impact on the type of feeding (Table 1) and the composition of diets, which has a certain effect on the microbiota of intestinal contents and methanogenic archaea in particular.Table 1 Animal diets in different regions of Kazakhstan.Full size tableIn the course of the research work, regions and specific agricultural formations were identified in the context of these regions.In North Kazakhstan, the fodder base is represented by such fodders as alfalfa hay, herb hay, alfalfa haylage, wheat straw, fodder wheat and sunflower cake. The feed is mainly of 2 quality classes. The live weight of cattle ranged from 375 to 480 kg. Feeding type: hay-concentrate and haylage-hay-concentrate.In the Western region, the animals were on the pasture, represented by the green mass of feather grass, hair, sage and tansy. Beef cattle are represented by the following breeds: Kazakh white-headed, Aberdeen-Angus and Hereford. Average live weight is 350–550 kg.In the Southeast region, the fodder base consists of wheat hay, sainfoin + alfalfa hay, mountain hay, herb haylage, corn silage and crushed corn. The feed is mainly of 2 and 3 classes. Hay-concentrate type of feeding is used, as well as pastures. Livestock of Angus, Kazakh white-headed breeds and animals of the local population are kept. Live weight of young animals is in the range of 360–380 kg.The diets of the Southern Region include natural grass hay, alfalfa hay, wheat straw, alfalfa haylage and concentrates. Hay-concentrate type of livestock feeding is widespread in the region. The average live weight of bulls for fattening of the Kazakh white-headed and Angus breeds—360–420 kg with a daily increase in live weight of 870–920 g.The composition of the fecal microbiota depending on the type of feeding is presented in Table 2.Table 2 The content of methanogenic archaea in feces.Full size tableFrom the data of Table 2 it follows that the largest amount of Bacteria was found in the faeces of animals with silage-concentrated feeding (98.59 ± 13.0%), and the smallest—with pasture-concentrated (93.24 ± 3.73%) and haylage—concentrated (93.8 ± 12.41%) types of feeding. The differences amounted to 5.35 and 4.79 absolute percent, respectively. However, the differences were not significant at P  More

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    Fractal features of soil grain-size distribution in a typical Tamarix cones in the Taklimakan Desert, China

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    Evolutionary implications of new Postopsyllidiidae from mid-Cretaceous amber from Myanmar and sternorrhynchan nymphal conservatism

    Systematic palaeontologyOrder Hemiptera Linnaeus, 1758Suborder Sternorrhyncha Amyot et Audinet-Serville, 1843Superfamily Protopsyllidioidea Carpenter, 1931Family Postopsyllidiidae Hakim, Azar et Huang, 2019Genus Megalophthallidion Drohojowska et Szwedo, gen. nov.LSID urn:lsid:zoobank.org:act:A6F71390-9B8E-4A19-8F30-C2A024B6EFB1Type speciesMegalophthallidion burmapateron Drohojowska et Szwedo, sp. nov.; by present designation and monotypy.EtymologyGeneric name is derived from Classic Greek megas (μέγας)—large, ophthalmos (ὀφθαλμός)—an eye and Greek form of generic name Psyllidium. Gender: masculine.Type localityNorthern Myanmar: state of Kachin, Noije bum 2001 Summit Site amber mine in the Hukawng Valley, SW of Maingkhwan.Type stratumLowermost Cenomanian, Upper Cretaceous (‘mid-Cretaceous’).DiagnosisHead capsule with 12 stiff setae on tubercles (18 setae in Postopsyllidium); fore wing without pterostigma (tiny pterostigma, widening of ScP + RA present in Postopsyllidium); vein CuP not thickened distally (distinctly thickened distally in Postopsyllidium); profemur with a row of ventral (ventrolateral) setae (two rows in Postopsyllidium).Megalophthallidion burmapateron Drohojowska et Szwedo, sp. nov.LSID urn:lsid:zoobank.org:act:F3F971F4-AE04-4F41-98B0-9A0A04470625.(Figs. 1A–F, 2A–I).Figure 1Megalophthallidion burmapteron gen. et sp. nov., holotype (MAIG 6687), imago. (A) Photo of body, ventral side; (B) photo of right antennae and (C) drawing of antenna; (D) drawing of body, dorsal side; (E) drawing of thorax structure with sclerites marked: red—pronotum; orange—mesopraescutum; yellow—mesoscutum; light green—mesoscutellum, dark green—mesopostnotum; light blue—metascutum; dark blue—metascutellum; violet—metapostnotum; (F) photo of thorax dorsal side. Scale bars: 0.5 mm (A), 0.2 mm (B–D), 0.1 mm (F).Full size imageFigure 2Megalophthallidion burmapteron gen. et sp. nov., holotype (MAIG 6687), imago. (A) Photo of right fore wing; (B) photo of right wings; (C) photo of antenna and proleg; (D) photo of proleg and mesoleg, and (E) photo of femur of proleg, and (F) photo of right metatarsus and left mesotarsus in the background, and (G) photo of right mesotarsus of mesoleg, and (H) Photo of tarsi; (I) photo of male genital block. Scale bars: 0.5 mm (A–D), 0.2 mm (B,E,F,H), 0.1 mm (G,I).Full size imageMaterialHolotype, number MAIG 6687 (BUB 96), deposited in Museum of Amber Inclusions (MAIG), University of Gdańsk, Poland. Imago, a complete and well-preserved male. Piece of amber 8 × 6 × 3 mm, cut from larger lump, polished flat on both sides.Type localityNorthern Myanmar: state of Kachin, Noije bum 2001 Summit Site amber mine in the Hukawng Valley, SW of Maingkhwan.Type stratumLowermost Cenomanian, Upper Cretaceous (‘mid-Cretaceous’).DiagnosisAs for the genus with the following additions: three ocelli distinct, antennomere IX the longest, about as long as pedicel, antennomeres III–VII and XI of similar length, antennomere XII the shortest, subconically tapered in apical portion. Paramere lobate, ventral margin with acute, small process, apical and dorsal margins rounded. Aedeagus geniculately bent at base, directed dorsally, tapered apicad.DescriptionMale (Figs. 1A–F, 2A–I). Head with compound eyes distinctly wider than pronotum (Fig. 1D–F). Compound eyes subglobular, protruding laterally. Vertex short in midline, about 2.5 times as wide as posterior margin and as long in middle; trapezoidal, anterior margin slightly arched, lateral margins diverging posteriad, posterior margin shallowly arched, disc of vertex with distinct setae on large tubercles: four setae at posterior margin, two at anterior angles of compound eyes, two medial, over the median ocellus. Three ocelli present, median ocellus distinct, visible from above, lateral ocelli near anterior angles of compound eyes. Frons about as wide as long in midline, two rows of setae on tubercles, upper row at level of median ocellus, lower one, below half of compound eye height. Clypeus, elongate, triangular, in lower portion roof-like; two setae on tubercles near upper margin. Genae very narrow. Rostrum reaching slightly beyond mesocoxae, apical segment slightly shorter than subapical one, darker. Antennae bases placed at lower margin of compound eyes; antennal fovea elevated; scapus shorter than pedicel, cylindrical; pedicel cylindrical; antennomeres IIIrd–VIIth and XIth of similar length, VIIIth slightly longer than VIIth, as long as Xth antennomere, IXth the longest, XIIth the shortest, tapered apically; rhinaria absent.Thorax (Fig. 1D–F): pronotum quadrangular, about as long as mesothorax; pronotum with anterior and posterior margins parallel, merely arcuate, disc with transverse groove in the median portion, lateral margins slightly arcuate, two distinct setae on tubercles in anterolateral angle, two setae on tubercles anterior margin at distance1/3 to median line, three distinct setae on tubercles in posterolateral angles. Mesopraescutum subtriangular, with apex widely rounded, about 0.4 times as wide as pronotum, about 0.4 times as long as wide, delicately separated from mesoscutum. Mesoscutum as wide as pronotum at widest point, distinctly narrowed medially, anterior angles rounded, anterolateral margin sigmoid, lateral angle acute, posterior angles wide, posterior margin V-shape incised, posterolateral areas of mesoscutum disc declivent posteriorly; disc with two setae on tubercles, at 1/3 of mesoscutum width. Mesoscutellum about as long as wide, diamond-shape, anterior and lateral angles acute, posterior angle rounded. Mesopostnotum in form of transverse band, slightly widened in median portion. Metascutum narrower than mesoscutum, anterior angles widely rounded, lateral angles acute, anterolateral margin concave, posterior margin arcuate, with deep median arcuate incision. The suture between metascutum and metascutellum weakly visible, metascutellum subtriangular, longer than wide at base.Parapteron with three distinct setae.Fore wing (Fig. 2A,B) membranous, narrow, elongate, about 3.5 times as long as wide, widest at 2/3 of length. Anterior margin merely arcuate, slightly bent at very base, anteroapical angle widely arcuate, apex rounded, posteroapical angle widely arcuate, tornus arcuate, claval margin straight, with incision between terminals of Pcu (claval apex) and A1. Stem ScP + R + MP + CuA slightly arcuate, very short stalk ScP + R + MP + CuA leaving basal cell, stem ScP + R oblique, straight, forked in basal half of fore wing length, branch ScP + RA, oblique, reaching anterior margin slightly distally of half of fore wing length, slightly distally of ending of CuA2 branch; branch RP slightly arcuate, a little more curved in basal section, reaching margin at anteroapical angle; stalk MP + CuA slightly shorter than basal cell; stem MP almost straight, forked in apical half of fore wing, at about 2/3 of fore wing length, with three terminals reaching margin between apex and posteroapical angle; stem CuA shorter than branches CuA1 and CuA2, about half as long as branch CuA1; claval vein CuP weak at base, not thickened distally; claval vein Pcu straight, claval vein A1 straight. Basal cell present, subtriangular, about twice as long as wide, basal veinlet cua-cup oblique, no other veinlets present; cell r (radial) very long, longer than half of fore wing length; cell m (medial) the shortest, shorter than cell cu (areola postica). Margins of fore wing with fringe of long setae, starting on costal margin near base of fore wing, ending at level of middle of cell cu; longitudinal veins with distinct, scarcely but evenly dispersed, movable setae; terminal section of CuP with two setae; costal margin with row of short, densely distributed setae, apical margin, tornus and claval margin with rows of scaly setae.Hind wing (Fig. 2B) membranous, shorter than fore wing, 3.23 times as long as wide. Costal margin bent at base, then almost straight up to the level of ScP + RA end and wing coupling lobe, then straight to anteroapical angle, anteroapical angle widely arcuate, apex arcuate, posteroapical angle arcuate, tornus straight, claval margin merely arcuate, posteroclaval angle angulate; stem ScP + R + MP bent at base, then straight, stem ScP + R short, branch ScP + RA short, about as long as stem ScP + R, branch RP arcuate basally than straight, reaching apex; stem MP arcuate, forked slightly distad CuA1 terminus level, branch MP1+2 slightly arcuate, reaching margin at posteroapical angle, branch MP3+4 straight, reaching tornus; stem CuA slightly bent at base, then straight, forked slightly distad ScP + R forking, branch CuA1 arcuate, branch CuA2 short, straight, slightly oblique, reaching tornus; claval vein CuP weak, visible only at base, claval vein Pcu slightly arcuate; wing coupling apparatus (fold) with a few short setae.Legs slender, relatively long, profemora armed (Fig. 2C–H). Procoxa as long as profemur, narrow, flattened. Protrochanter scaphoid, elongate, with long apical and subapical setae. Profemur flattened laterally, about as long as protibia, ventrally armed with four large setae on elevated plinths; dorsal margin with row of short, decumbent setae. Protibia narrow, rounded in cross section, covered with short setae, a few longer setae in distal portion. Protarsus—single, long tarsomere, plantar surface with row of semi-erect setae; tarsal claws long, straight, directed ventrally, no arolium nor empodium.Mesocoxa elongate, narrow, slightly flattened. Mesotrochanter scaphoid. Mesofemur slender, flattened laterally, dorsal margin with short setae. Mesotibia subequal to mesofemur, slender, covered with setae, two apical setae slightly thicker and longer. Mesotarsus with three tarsomeres, basimesotarsomere the longest, shorter than cumulative length of mid- and apical mesotarsomere, plantar margins with setae, two apical setae slightly longer and thicker; midmesotarsomere the shortest, 1/3 of basimesotarsomere length, a few setae on plantar surface; apical tarsomere shorter than basimesotarsomere, twice as long as midmesotarsomere, plantar surface with a few, scarcely dispersed setae, tarsal claws long, narrow, directed ventrally, no arolium nor empodium.Metacoxa conical, narrow. Metatrochanter scaphoid, elongate. Metafemur slender, laterally flattened, longer than mesofemur, dorsal margin with row of short setae. Metatibia, long, slender, 1.6 times as long as metafemur, with suberect setae of different size, two larger and longer and two shorter setae subapical setae. Metatarsus slightly less than half of metatibia length, with three tarsomeres, basimetatarsomere the longest, more than twice as long as apical metatarsomere, 1.5 times as long as combined length of mid- and apical metatarsomere, plantar surface with scarce decumbent setae; mid metatarsomere the shortest, 1/4 of basimetatarsomere length, plantar surface with a few setae, two apical ones slightly thicker; apical metatarsomere about 0.4 of basimetatarsomere length, with scarcely dispersed setae on along plantar surface; tarsal claws, long, slender, other pretarsal structures absent.Abdomen (Fig. 1F) narrowly attached to thorax, tergite segment shorter, 2nd tergite distinctly longer, 3rd to 8th tergites of similar length; pygofer narrowing apicad, ventral margin strongly elongated posteriorly; anal tube short, directed posterodorsad, anal style shorter than anal tube. Paramere lobate, ventral margin with acute, small process, apical and dorsal margins rounded. Aedeagus (Fig. 2I) geniculately bent at base, directed dorsad, tapered apicad.Female. Unknown.Megalophthallidion sp. (5th instar nymph)(Figs. 3A–D, 4A–F)Figure 3Megalophthallidion sp. (MAIG 6688), nymph. (A) Photo of body, dorsal side and (B) drawing of body dorsal side; (C) photo of body dorsal side and (D) drawing of body ventral side. Scale bars: 0.5 mm (A–D).Full size imageFigure 4Megalophthallidion sp. (MAIG 6688), nymph. Photo of clypeus and (B) drawing of clypeus; (C) photo of proleg, and (D) photo of mesoleg, and (E) photo of metaleg; (F) photo of posterior part of abdomen ventral side. Scale bars: 0.1 mm (A–F).Full size imageMaterialNymph, 5th instar, MAIG 6688 (BUB 1799), deposited in Museum of Amber Inclusions (MAIG), University of Gdańsk, Poland. Piece of amber 13 × 6 × 2 mm, cut from larger lump, polished flat on one side, more convex on the other.Diagnostic charactersThe nymph of Megalophthallidion gen. nov. is similar in general body shape to the only known fossil protopsyllidioidean nymph described from Lower Cretaceous Lebanese amber—Talaya batraba Drohojowska et Szwedo, 2013. The nymph of Talaya batraba is 2nd or 3rd instar, therefore some features are difficult to compare with this last instar nymph of Megalophthallidion gen. nov. The morphological states observed in those two specimens are: head covered with strongly expanded disc and expanded disc of pronotum, however shapes and ratios of these structures differ; compound eyes on ventral side of head, shifted laterad (ommatidia on cones in T. batraba, while ventroposterior expansions are present in Megalophthallidion gen. nov.); compound eyes visible from above as short, stout cones in fissure between posterior margin of disc (hypertrophied vertex) and anterior margin of pronotum (compound eyes (?) are visible on dorsal side of Permian Aleuronympha bibulla Riek, 1974); in Megalophthallidion gen. nov. rostrum reached mesocoxa, while in Talaya batraba distinctly exceeds length of the body; abdomen with 9 segments; tergites of abdominal segments 5th–9th expanded posterolaterad in form of fan-like expansion; 9th abdominal segment short, placed ventral; anal tube short, cylindrical, epiproct (?) globular.DescriptionNymph, 5th instar (Figs. 3A–D, 4A–F). Body oval shaped, dorso-ventrally flattened, 1.5 times longer than wide with segmentation visible; on the ventral side slightly concave. Length of body c. 1.56 mm long, outline, in dorsal view, maximum width of body 0.94 mm; length of head and pronotum (cephaloprothorax) c. 0.46 mm in midline, width c. 0.83 mm; cumulative length of mesonotum + metanotum c. 0.25 mm; abdomen c. 0.8 mm long. Dorsal side (Fig. 3A,B) with distinct median line (ecdysial line), not reaching anterior or posterior margin of the body, the line distinctly roof-like in abdominal portion. Anterior margin of head (cephaloprothorax) disc arcuate, lateral angles rounded; anterior margin of pronotum arcuate, lateral margins arcuately diverging posteriad, posterior margin distinctly arcuate, anterior angles widely rounded, posterior angles acutely rounded, disc elevated, convex, lateral portions declivitous; the fissure between posterior margin of head disc and anterior margin of pronotum narrow, widened medially, with stalked compound eyes popping out.Head partly separated from prothorax, wide in ventral view. Bases of antennae protruding anterolaterally, wide, anterior margin arcuate, with a small lump extending anteriorly connecting margin with vertex expansion. Suture separating anteclypeus and postclypeus visible in ventral aspect (Fig. 4A,B). Postclypeus about three times as long as wide, oval, slightly swollen, without any setae; weak traces of salivary pump muscle attachments visible. Anteclypeus about as long as postclypeus, widened in upper section below clypeal suture, convex, carinately elevated in lower section, with sides distinctly declivitous, clypellus long, carinately elevated. Lora (mandibulary plates) distinct, separated from anteclypeus by shallow suture, with upper angles at half of postclypeus length, lower angles at half of anteclypeus length, about as wide as half of postclypeus width. Maxillary plates narrow. Genal portion of head enlarged, medial portion arcuately convex; lateral sections narrowing laterally, terminally encircling bases of compound eyes. Antennae short (Fig. 3C,D), placed in front of genal portion. Antennal flagellum indistinctly subdivided into four segments. Rostrum (Fig. 4A,B) three-segmented, 0.2 mm long, with apex reaching apex of mesocoxae; apical segment about 2.5 times as long as subapical one.No lateral sclerites on meso- and metathorax, only one plus one large medial sclerite on both meso- and metathorax. Mesothoracic and metathoracic wing pads distinct, wide, subtriangular, with posterior apices directed posteriorly; lateral portions of mesothoracic wing pads arcuate. Fore wing pad 0.6 mm long, with small, straight humeral lobe, forming a right angle, not protruding anteriorly. Mesothoracic tergites slightly larger than metathoracic segments (respectively c. 0.14 mm and c. 0.12 mm long in midline, 0.26 mm and 0.27 mm in lateral lines); mesothoracic tergum with distinct median elevation (low double crest with ecdysial line in between), slightly wider than long in midline, anterior margin arcuate, lateral margins straight, subparallel, posterior margin concave. Metathoracic wing pad apex slightly exceeding mesothoracic wing pad. Metathoracic tergum wider than long, slightly shorter than mesothoracic tergum, with distinct elevation in the middle.Legs relatively long (Figs. 3C,D, 4C–E). Coxae of legs placed near the median axis of the body. Prolegs: procoxal pit with margins elevated, procoxa conical (c. 0.1 mm long), protrochanter scaphoid, about as long as procoxa, profemur c. 0.13 mm long, slightly flattened laterally, merely thickened, protibia longer than profemur, c. 0.23 mm long; tarsus shorter than protibia, basiprotarsomere about as long as apical protarsomere, the latter with distinct tarsal claws, and wide arolium. Mesoleg similar to proleg, mesocoxa conical (c. 0.1 mm long), mesotrochanter scaphoid, mesofemur (c. 0.13 mm) slightly flattened laterally, mesotibia slightly longer than mesofemur (c. 0.18 mm), mesotarsus slightly shorter than mesotibia, three-segmented, basimesotarsomere the longest (c. 0.07 mm), about as long as combined length of mid- and apical mesotarsomeres (c. 0.04 mm respectively), arolium wide, tarsal claws distinct. Metaleg: metacoxa conical (c. 0.1 mm), metatrochanter scaphoid, about as long as metacoxa (c. 0.12 mm). Metafemur (c. 0.17 mm) slightly more thickened than pro- and mesofemur, metatibia slightly longer (0.19 mm) than pro- and mesotibiae. Metatarsus three-segmented: basimetatarsomere about as long (0.08 mm) as combined length of mid- and apical metatarsomeres (0.04 mm respectively), arolium lobate, wide, tarsal claws distinct, widely spread.Abdomen (Fig. 3A–D) 9-segmented, narrow at base, widening fan-shape posteriorly, 1st segment visible from above, segmentation visible, abdominal terga 5th–9th expanded posterolaterally. Tergites carinately elevated in the middle, separated by ecdysial line. 1st sternite visible in ventral view, sternites 2nd–4th fused medially, sternites 5th–9th separated; 9th abdominal segment short (Fig. 4F), placed ventrally, under tergal expansion; anal tube short, cylindrical, epiproct (?) globular. More