Dengler, J. et al. Biodiversity of palaearctic grasslands: A synthesis. Agric. Ecosyst. Environ. 182(1), 1–14 (2014).
Google Scholar
Wang, H. et al. Alpine grassland plants grow earlier and faster but biomass remains unchanged over 35 years of climate change. Ecol. Lett. 23, 701–710 (2020).
Google Scholar
Torok, P. et al. Step(pe) up! Raising the profile of the Palaearctic natural grasslands. Biodivers. Conserv. 25(12), 2187–2195 (2016).
Google Scholar
Kuss, F. R. & Graefe, A. R. Effects of recreation trampling on natural area vegetation. J. Leis. Res. 17, 165–183 (1985).
Google Scholar
Buckley, R. C. & Pannell, J. Environmental impacts of tourism and recreation in national parks and conservation reserves. J. Tourism Stud. 1, 24–32 (1990).
Yorks, T. et al. Toleration of traffic by vegetation: Life form conclusions and summary extracts from a comprehensive data base. Environ. Manage. 21, 12–131 (1997).
Google Scholar
Gouvenain, R. C. Indirect impacts of soil trampling on tree growth and plant succession in the north cascade mountains of Washington (1996).
Xu, L., Yu, F. H. & Drunen, E. V. Trampling, defoliation and physiological integration affect growth, morphological and mechanical properties of a root-suckering clonal tree. Ann. Bot. 109, 1001–1008 (2012).
Google Scholar
Bayfield, N. G. Use and deterioration of some Scottish hill paths. J. Appl. Ecol. 10, 639–648 (1973).
Google Scholar
Liddle, M. J. A selective review of the ecological effects of human trampling on natural ecosystems. Biol. Cons. 7, 17–36 (1975).
Google Scholar
Frissell, S. S. Judging recreational impacts on wilderness campsites. J. Forest. 76, 481–483 (1978).
Wagar, J. A. How to predict which vegetated areas will stand up best under ‘active’ recreation. Am. Recreat. J. 1, 20–21 (1961).
Cole, D. N. & Bayfield, N. G. Recreational trampling of vegetation: Standard experimental procedures. Biol. Cons. 63(3), 209–215 (1993).
Google Scholar
Prescott, O. & Stewart, G. Assessing the impacts of human trampling on vegetation: A systematic review and meta-analysis of experimental evidence. PeerJ 2, e360 (2014).
Google Scholar
Loreau, M. et al. Biodiversity and ecosystem functioning: Current knowledge and future challenges. Science 294, 804–808 (2001).
Google Scholar
Vandewalle, M. et al. Functional traits as indicators of biodiversity response to land use changes across ecosystems and organisms. Biodivers. Conserv. 19, 2921–2947 (2010).
Google Scholar
Petchey, O. L. & Gaston, K. J. Functional diversity, species richness and composition. Ecol. Lett. 5, 402–411 (2002).
Google Scholar
Cadotte, M. W., Carscadden, K. & Mirotchnick, N. Beyond species: Functional diversity and the maintenance of ecological processes and services. J. Appl. Ecol. 48(5), 1079–1087 (2011).
Google Scholar
Mouillot, D. et al. A functional approach reveals community responses to disturbances. Trends Ecol. Evol. 28, 167–177 (2013).
Google Scholar
Carmona, C. P. et al. Traits without borders: Integrating functional diversity across scales. Trends Ecol. Evol. 31(5), 382–394 (2016).
Google Scholar
Conradi, T. et al. Impacts of visitor trampling on the taxonomic and functional community structure of calcareous grassland. Appl. Veg. Sci. 18, 359–367 (2015).
Google Scholar
Pickering, C. M. & Barros, A. Using functional traits to assess the resistance of subalpine grassland to trampling by mountain biking and hiking. J. Environ. Manage. 164, 129–136 (2015).
Google Scholar
Baraloto, C., Herault, B. & Paine, C. E. T. Contrasting taxonomic and functional responses of a tropic tree community to selective logging. J. Appl. Ecol. 49, 861–870 (2012).
Google Scholar
Magnago, L. F. S. et al. Functional attributes change but functional richness is unchanged after fragmentation of Brazilian Atlantic forests. J. Ecol. 102, 475–485 (2014).
Google Scholar
Marion, J. L. & Cole, D. N. Spatial and temporal variation in soil and vegetation impacts on campsites. Ecol. Appl. 6, 520–530 (1996).
Google Scholar
Roovers, P. et al. Experimental trampling and vegetation recovery in some forest and heathland communities. Appl. Veg. Sci. 7, 111–118 (2004).
Google Scholar
Conradi, T. et al. Impacts of visitor trampling on the taxonomic and functional community structure of calcareous grassland. Appl. Veg. Sci. 18(3), 359–367 (2015).
Google Scholar
Zamora, R. Functional equivalence in plant-animal interactions: Ecological and evolutionary consequences. Oikos 88(2), 442–447 (2000).
Google Scholar
Hubbell, S. P. Neutral theory in community ecology and the hypothesis of functional equivalence. Funct. Ecol. 19, 166–172 (2005).
Google Scholar
Mouchet, M. A. et al. Functional diversity measures: An overview of their redundancy and their ability to discriminate community assembly rules. Funct. Ecol. 24, 867–876 (2010).
Google Scholar
Diaz, S. et al. Incorporating plant functional diversity effects in ecosystem service assessments. Proc. Natl. Acad. Sci. U.S.A. 104, 20684–20689 (2007).
Google Scholar
Suding, K. N. & Goldstein, L. J. Testing the Holy Grail framework: Using functional traits to predict ecosystem change. New Phytol. 180, 559–562 (2008).
Google Scholar
De Bello, F. et al. Towards an assessment of multiple ecosystem processes and services via functional traits. Biodivers. Conserv. 19, 2873–2893 (2010).
Google Scholar
Devictor, V. et al. Spatial mismatch and congruence between taxonomic, phylogenetic and functional diversity: The need for integrative conservation strategies in a changing world. Ecol. Lett. 13, 1030–1040 (2010).
Google Scholar
Tian, K., Guo, H. J. & Yang, Y. M. Ecological structures and functions of plateau wetlands in China (Chinese Science Press, 2009).
Garnier, E. et al. standardized protocol for the determination of specific leaf size and leaf dry matter content. Funct. Ecol. 15, 688–695 (2001).
Google Scholar
Cornelissen, J. H. C. et al. A handbook of protocols for standardised and easy measurement of plant functional traits worldwide. Aust. J. Bot. 51, 335–380 (2003).
Google Scholar
Mason, N. W. H. et al. An index of functional diversity. J. Veg. Sci. 14, 571–578 (2003).
Google Scholar
Villeger, S., Mason, N. W. H. & Mouillot, D. New multidimensional functional diversity indices for a multifaceted framework in functional ecology. Ecology 89, 2290–2301 (2008).
Google Scholar
Lavorel, S. & Garnier, E. Predicting changes in community composition and ecosystem functioning from plant traits: Revisiting the Holy Grail. Funct. Ecol. 16, 545–556 (2002).
Google Scholar
Cianciaruso, M. V. et al. Including intraspecific variability in functional diversity. Ecology 90, 81–89 (2009).
Google Scholar
Lepš, J. et al. Community trait response to environment: Disentangling species turnover vs intraspecific trait variability effects. Ecography 34, 856–863 (2011).
Google Scholar
Garnier, E. et al. Assessing the effects of land-use change on plant traits, communities and ecosystem functioning in grasslands: A standardized methodology and lessons from an application to 11 European sites. Ann. Bot. 99, 967–985 (2007).
Google Scholar
Pakeman, R. J. & Quested, H. M. Sampling plant functional traits: What proportion of the species need to be measured?. Appl. Veg. Sci. 10, 91–96 (2007).
Google Scholar
Hothorn, T., Bretz, F. & Westfall, P. Simultaneous inference in general parametric models. Biom. J. 50, 346–363 (2008).
Google Scholar
Oksanen, J., Blanchet, F. G., Friendly, M. et al. Vegan: Community Ecology Package. R package version 2.5–7 (2020).
Laliberté, E., Legendre, P., Shipley, B. FD: measuring functional diversity from multiple traits, and other tools for functional ecology. R package version 1.0–12 (2014).
